Bivalvia | Unionida | Unionoidea | Unionidae | Parreysiinae

tribe INDOCHINELLINI Bolotov, Pfeiffer, Vikhrev, & Konopleva in Bolotov et al., 2018: cladomics

9 genera

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 34 (34 total)
number of subtaxa (rank: genus) = 9
analyzed in 69 cladograms in 43 publications
maximum number of terminals in an analysis = 31 (mean = 9.35, median = 4)
maximum number of subtaxa (genus) = 8 (mean = 3.43, median = 3)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 42
maximum number of included terminals = 31, (mean = 14.71 median = 12)
maximum number of subtaxa (genus) = 8 (mean = 5, median = 4)
proportion of cladograms that support basic monophyly = 0.95

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 38
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (genus) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 31 (mean = 15.47, median = 12)
maximum number of subtaxa (genus) = 8 (mean = 5.18, median = 4)
proportion of cladograms that support rigorous monophyly = 0.95


Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Indochinellini cladogram distances
PCoA Ordination

NJ Tree of Indochinellini cladogram distances
NJ Tree


Cladograms

Bolotov et al., 2024 [Publ 4369], fig. 2

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

cstiTtTbiriSoTroTroSiD
0.957 29 29 27 8 33 33 4 **
LAMELLIDENTINI LEOPARREYSIINI PARREYSIINI COELATURINI Pseudoscabies Indonaia Indochinella Radiatula Scabiellus Harmandia Unionetta Scabies

Bolotov et al., 2024 [Publ 4369], fig. S1

cstiTtTbiriSoTroTroSiD
0.913 29 29 27 8 34 33 4 16.0
Parreysia LEOPARREYSIINI LAMELLIDENTINI Assamnaia COELATURINI Pseudoscabies Indonaia Indochinella Radiatula Scabiellus Harmandia Unionetta Scabies

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

cstiTtTbiriSoTroTroSiD
0.741 29 29 28 8 604 56 4 21.0
PARREYSIINI COELATURINI Pseudoscabies Indonaia Indochinella Radiatula Scabiellus Unionetta fabagina 2 Harmandia Unionetta fabagina 1 Scabies

Konopleva et al., 2026 [Publ 4847], fig. 2

cstiTtTbiriSoTroTroSiD
0.957 31 31 28 8 35 35 4 19.0
LAMELLIDENTINI LEOPARREYSIINI PARREYSIINI COELATURINI Pseudoscabies Indonaia Indochinella Radiatula Scabiellus Scabies Unionetta Harmandia

Konopleva et al., 2026 [Publ 4847], fig. S1

cstiTtTbiriSoTroTroSiD
0.957 31 31 29 8 35 35 4 8.0
LAMELLIDENTINI LEOPARREYSIINI PARREYSIINI COELATURINI Pseudoscabies Indonaia Indochinella Radiatula Scabiellus Harmandia Unionetta Scabies

Konopleva et al., 2026 [Publ 4847], fig. S2

cstiTtTbiriSoTroTroSiD
0.913 31 31 29 8 36 35 4 16.0
Parreysia LEOPARREYSIINI LAMELLIDENTINI Assamnaia COELATURINI Pseudoscabies Indonaia Indochinella Radiatula Scabiellus Harmandia Unionetta Scabies

Konopleva et al., 2026 [Publ 4847], fig. S3

cstiTtTbiriSoTroTroSiD
0.913 31 31 26 8 36 35 4 15.0
Parreysia Assamnaia LEOPARREYSIINI LAMELLIDENTINI COELATURINI Pseudoscabies Indonaia Indochinella Radiatula Scabiellus Harmandia Unionetta Scabies

Pfeiffer et al., 2018 [Publ 3827], fig. 2

cstiTtTbiriSoTroTroSiD
0.950 17 17 15 7 11 10 4 16.0
LAMELLIDENTINI PARREYSIINI LEOPARREYSIINI COELATURINI Indonaia Indochinella Radiatula Scabiellus Unionetta fabagina 2 Harmandia Unionetta fabagina 1 Scabies

Jeratthitikul et al., 2019 [Publ 3922], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 18 18 16 7 5 5 4 21.0
LAMELLIDENTINI LEOPARREYSIINI PARREYSIINI COELATURINI Indonaia Indochinella Radiatula Scabiellus Harmandia Unionetta Scabies

Bolotov et al., 2020 [Publ 4100], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 21 21 19 7 96 20 4 18.0
LAMELLIDENTINI PARREYSIINI LEOPARREYSIINI COELATURINI Indonaia Indochinella Radiatula Scabiellus Scabies Unionetta Harmandia

Bolotov et al., 2020 [Publ 4100], fig. S1

cstiTtTbiriSoTroTroSiD
0.947 20 20 17 7 91 20 4 16.0
LAMELLIDENTINI PARREYSIINI LEOPARREYSIINI COELATURINI Indonaia Indochinella Radiatula mouhoti Radiatula myitkyinae Radiatula chaudhurii Radiatula crispisulcata Scabiellus Harmandia Unionetta Scabies

Bolotov et al., 2022 [Publ 4239], fig. 2

cstiTtTbiriSoTroTroSiD
0.950 26 26 24 7 32 28 4 17.0
LAMELLIDENTINI PARREYSIINI LEOPARREYSIINI COELATURINI Indonaia Indochinella Radiatula Scabiellus Harmandia Unionetta Scabies

Bolotov et al., 2022 [Publ 4239], fig. S1

cstiTtTbiriSoTroTroSiD
0.950 26 26 23 7 32 28 4 4.0
LAMELLIDENTINI PARREYSIINI LEOPARREYSIINI COELATURINI Indonaia Indochinella Radiatula Scabiellus Harmandia Unionetta Scabies

Bolotov et al., 2022 [Publ 4239], fig. S2

cstiTtTbiriSoTroTroSiD
0.950 26 26 24 7 32 28 4 21.0
PARREYSIINI LAMELLIDENTINI LEOPARREYSIINI COELATURINI Indonaia Indochinella Radiatula Scabiellus Harmandia Unionetta Scabies

Bolotov et al., 2022 [Publ 4252], fig. 2

cstiTtTbiriSoTroTroSiD
0.950 27 27 25 7 34 30 4 2.0
LAMELLIDENTINI PARREYSIINI LEOPARREYSIINI COELATURINI Indonaia Indochinella Radiatula Scabiellus Harmandia Unionetta Scabies

Bolotov et al., 2022 [Publ 4252], fig. S1

cstiTtTbiriSoTroTroSiD
0.950 27 27 25 7 34 30 4 22.0
PARREYSIINI LAMELLIDENTINI LEOPARREYSIINI COELATURINI Indonaia Indochinella Radiatula Scabiellus Harmandia Unionetta Scabies

Chen et al., 2026 [Publ 4833], fig. 1

cstiTtTbiriSoTroTroSiD
0.800 14 15 14 7 13 10 4 29.0
Lamellidens Trapezidens PARREYSIINI LEOPARREYSIINI COELATURINI Indochinella Indonaia Radiatula PSEUDODONTINI Harmandia Chenunionetta Unionetta fabagina 2 Unionetta fabagina 1 Scabies

Konopleva et al., 2023 [Publ 4262], fig. 8

cstiTtTbiriSoTroTroSiD
1.000 14 14 10 7 68 0 0 23.0
PSEUDODONTINI RECTIDENTINI GONIDEINI LAMPROTULINI Indonaia Indochinella Radiatula Scabiellus Scabies Unionetta Harmandia

Bolotov et al., 2017 [Publ 3761], fig. S1

cstiTtTbiriSoTroTroSiD
0.882 11 11 9 4 128 18 4 28.0
LAMELLIDENTINI PARREYSIINI LEOPARREYSIINI COELATURINI Indonaia Indochinella Scabies crispata Radiatula Scabies mandarinus Scabies humilis Scabies phaselus

Bolotov et al., 2017 [Publ 3761], fig. S2

cstiTtTbiriSoTroTroSiD
0.882 11 11 9 4 128 18 4 28.0
PARREYSIINI LAMELLIDENTINI LEOPARREYSIINI COELATURINI Indonaia Indochinella Scabies crispata Radiatula Scabies mandarinus Scabies humilis Scabies phaselus

Bolotov et al., 2017 [Publ 3764], fig. S1

cstiTtTbiriSoTroTroSiD
0.647 12 13 9 4 113 26 4 24.0
Trapezidens Lamellidens lamellatus Lamellidens jenkinsianus Lamellidens generosus Lamellidens mainwaringi Lamellidens ferrugineus Lamellidens marginalis Lamellidens savadiensis Lamellidens corrianus 1 PARREYSIINI LEOPARREYSIINI COELATURINI Indochinella Scabies Radiatula Indonaia shurtleffiana Indonaia rugosa Indonaia andersoniana Indonaia caerulea Indonaia bonneaudii Lamellidens corrianus 2

Bolotov et al., 2018 [Publ 3826], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 11 11 9 4 71 13 4 24.0
LAMELLIDENTINI PARREYSIINI LEOPARREYSIINI COELATURINI Indonaia Indochinella Radiatula Scabies

Huang et al., 2019 [Publ 3843], fig. 2

cstiTtTbiriSoTroTroSiD
0.941 12 12 9 4 189 22 4 26.0
LAMELLIDENTINI PARREYSIINI LEOPARREYSIINI COELATURINI Indonaia Indochinella Scabies Radiatula

Huang et al., 2019 [Publ 3843], fig. S3

cstiTtTbiriSoTroTroSiD
0.941 12 12 9 4 187 22 4 24.0
LAMELLIDENTINI PARREYSIINI LEOPARREYSIINI COELATURINI Indonaia Indochinella Scabies Radiatula

Huang et al., 2019 [Publ 3843], fig. S4

cstiTtTbiriSoTroTroSiD
0.941 12 12 9 4 189 22 4 24.0
PARREYSIINI LAMELLIDENTINI LEOPARREYSIINI COELATURINI Indonaia Indochinella Scabies Radiatula

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

cstiTtTbiriSoTroTroSiD
1.000 4 4 2 4 53 6 3 25.0
LAMELLIDENTINI LEOPARREYSIINI COELATURINI Indonaia Indochinella Scabies Harmandia

Pfeiffer et al., 2019 [Publ 3937], fig. 6

cstiTtTbiriSoTroTroSiD
1.000 4 4 2 4 53 6 3 25.0
LAMELLIDENTINI LEOPARREYSIINI COELATURINI Indonaia Indochinella Scabies Harmandia

Ortiz-Sepulveda et al., 2020 [Publ 4101], fig. 1

cstiTtTbiriSoTroTroSiD
0.812 8 8 3 4 28 28 2 24.0
LEOPARREYSIINI COELATURINI Indonaia Indochinella Radiatula mouhoti Radiatula myitkyinae Radiatula chaudhurii Scabies

Ortiz-Sepulveda et al., 2020 [Publ 4101], fig. 2

cstiTtTbiriSoTroTroSiD
0.812 8 8 1 4 28 28 2 26.0
LEOPARREYSIINI COELATURINI Radiatula mouhoti Radiatula myitkyinae Radiatula chaudhurii Indochinella Indonaia Scabies

Konopleva et al., 2019 [Publ 3900], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 12 12 9 4 39 0 0 22.0
RECTIDENTINI Indonaia Indochinella Scabies Radiatula

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 3 3 1 3 37 5 3 26.0
LAMELLIDENTINI LEOPARREYSIINI COELATURINI Indochinella Radiatula Scabies

Konopleva et al., 2017 [Publ 3807], fig. 6

cstiTtTbiriSoTroTroSiD
1.000 3 3 1 3 28 8 3 26.0
LAMELLIDENTINI LEOPARREYSIINI COELATURINI Indochinella Radiatula Scabies

Wu et al., 2021 [Publ 4366], fig. 2_I

cstiTtTbiriSoTroTroSiD
1.000 5 5 3 3 86 8 3 26.0
LAMELLIDENTINI LEOPARREYSIINI COELATURINI Indonaia Scabies Radiatula

Wu et al., 2021 [Publ 4366], fig. 6

cstiTtTbiriSoTroTroSiD
0.917 5 5 3 3 86 8 3 28.0
LAMELLIDENTINI LEOPARREYSIINI COELATURINI Indonaia Radiatula myitkyinae Scabies Radiatula mouhoti Radiatula chaudhurii

Bolotov et al., 2022 [Publ 4250], fig. 8

cstiTtTbiriSoTroTroSiD
1.000 9 9 7 3 40 15 3 22.0
LAMELLIDENTINI PARREYSIINI LEOPARREYSIINI Indonaia Indochinella Radiatula

Konopleva et al., 2023 [Publ 4269], fig. S1_4

cstiTtTbiriSoTroTroSiD
1.000 3 3 1 3 53 4 3 26.0
LAMELLIDENTINI PARREYSIINI LEOPARREYSIINI Indonaia Indochinella Radiatula

Bolotov et al., 2022 [Publ 4250], fig. S2

cstiTtTbiriSoTroTroSiD
1.000 12 12 10 3 3 3 2 21.0
PARREYSIINI LEOPARREYSIINI Indonaia Indochinella Radiatula

Whelan et al., 2011 [Publ 3380], fig. 1-ML

cstiTtTbiriSoTroTroSiD
0.923 2 2 0 2 34 7 3 --
LAMELLIDENTINI LEOPARREYSIINI COELATURINI Radiatula Indochinella

Whelan et al., 2011 [Publ 3380], fig. 1-MP

cstiTtTbiriSoTroTroSiD
0.923 2 2 0 2 34 7 3 --
LAMELLIDENTINI LEOPARREYSIINI COELATURINI Radiatula Indochinella

Bolotov et al., 2019 [Publ 3974], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 6 6 4 2 12 7 2 23.0
LEOPARREYSIINI COELATURINI Indochinella Radiatula

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 80 3 3 --
LAMELLIDENTINI LEOPARREYSIINI COELATURINI Indochinella

Zieritz et al., 2024 [Publ 4381], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 44 4 3 --
LAMELLIDENTINI LEOPARREYSIINI Unionetta COELATURINI

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 121 2 2 --
COELATURINI LEOPARREYSIINI Radiatula

Choi et al., 2020 [Publ 4169], fig. 5

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 9 2 2 --
Radiatula LAMELLIDENTINI LEOPARREYSIINI

Chen et al., 2023 [Publ 4302], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 40 2 2 --
COELATURINI Indonaia PARREYSIINI

Chen et al., 2023 [Publ 4302], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 40 2 2 --
PARREYSIINI COELATURINI Indonaia

Imsong & Murali, 2023 [Publ 4370], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 2 2 2 --
LAMELLIDENTINI Indonaia PARREYSIINI

Dai et al., 2025 [Publ 4440], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 43 2 2 --
PARREYSIINI COELATURINI Indonaia

Dai et al., 2025 [Publ 4440], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 43 2 2 --
COELATURINI Indonaia PARREYSIINI

Chen et al., 2025 [Publ 4487], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 40 2 2 --
PARREYSIINI COELATURINI Indonaia

Chen et al., 2025 [Publ 4487], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 40 2 2 --
PARREYSIINI COELATURINI Indonaia

Dai et al., 2024 [Publ 4385], fig. 3-BI

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 30 1 1 --
LAMELLIDENTINI Scabies

Dai et al., 2024 [Publ 4385], fig. 3-ML

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 30 1 1 --
LAMELLIDENTINI Scabies

Dai et al., 2024 [Publ 4386], fig. 5-BI

cstiTtTbiriSoTroTroSiD
1.000 2 2 0 1 20 1 1 --
LAMELLIDENTINI Scabies

Dai et al., 2024 [Publ 4386], fig. 5-ML

cstiTtTbiriSoTroTroSiD
1.000 2 2 0 1 20 1 1 --
LAMELLIDENTINI Scabies

Dai et al., 2026 [Publ 4757], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 33 1 1 --
Scabies LAMELLIDENTINI

Combosch et al., 2017 [Publ 3806], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 14 0 0 --
LAMPSILINI Scabies

Combosch et al., 2017 [Publ 3806], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 14 0 0 --
Scabies LAMPSILINI

Huang et al., 2018 [Publ 3828], fig. 3_5

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 28 0 0 --
LAMPSILINI Scabies

Jeratthitikul et al., 2019 [Publ 3911], fig. 4

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 8 0 0 --
PSEUDODONTINI Scabies

Wu et al., 2020 [Publ 4121], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 33 0 0 --
LAMPSILINI Scabies

Wu et al., 2020 [Publ 4121], fig. S1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 33 0 0 --
LAMPSILINI Scabies

Sano et al., 2020 [Publ 4138], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 68 0 0 --
Scabies LAMPSILINI CHAMBERLAINIINI Inversidens LAMPROTULINI Ptychorhynchus Obovalis Anemina Buldowskia Pletholophus Beringiana Sinanodonta Cristaria ANODONTINA ALASMIDONTINA LANCEOLARIINI LEPIDODESMINI UNIONINI

Sano et al., 2020 [Publ 4138], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 68 0 0 --
Scabies CHAMBERLAINIINI LAMPSILINI LAMPROTULINI GONIDEINI Anemina Buldowskia Sinanodonta Pletholophus Beringiana Cristaria ANODONTINA ALASMIDONTINA LANCEOLARIINI LEPIDODESMINI UNIONINI

Sano et al., 2020 [Publ 4138], fig. 4

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 68 0 0 --
Scabies LAMPSILINI CHAMBERLAINIINI Inversidens LAMPROTULINI Obovalis Ptychorhynchus Anemina Buldowskia Sinanodonta Pletholophus Beringiana Cristaria ANODONTINA ALASMIDONTINA LEPIDODESMINI LANCEOLARIINI UNIONINI

Wu et al., 2021 [Publ 4215], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 23 0 0 --
Indonaia ACUTICOSTINI UNIONINI CHAMBERLAINIINI PSEUDODONTINI Sinosolenaia LAMPROTULINI Ptychorhynchus Parvasolenaia Inversidens Gonidea Microcondylaea Leguminaia

Smith, 2021 [Publ 4331], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 19 0 0 --
Scabies ANODONTINI

Dai et al., 2024 [Publ 4386], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 13 0 0 --
Scabies UNIONINA MIDDENDORFFINAIINA OXYNAIINA ACULAMPROTULINA ANODONTINI LEPIDODESMINI

Xiang et al., 2025 [Publ 4433], fig. 6

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 27 0 0 --
Scabies ANODONTINI