Bivalvia | Unionida | Unionoidea | Margaritiferidae | Margaritiferinae

genus Pseudunio Haas, 1910: cladomics

3 Recent valid species (5 total species)

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 3 (5 total)
analyzed in 81 cladograms in 44 publications
maximum number of terminals in an analysis = 5 (mean = 1.53, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 31
maximum number of included terminals = 5, (mean = 2.39 median = 2)
proportion of cladograms that support basic monophyly = 0.94


Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Pseudunio cladogram distances
PCoA Ordination

NJ Tree of Pseudunio cladogram distances
NJ Tree


Cladograms

Lyubas et al., 2019 [Publ 3963], fig. 10

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
5 5 1 5 11 8 2 0
  +----------- Margaritifera             
  |                                      
--+  +-------- Cumberlandia              
  |  |                                   
  +--+  +----- Pseudunio_marocanus **       
     |  |                                
     +--+  +-- Pseudunio_auricularius **    
        |  |                             
        |  +-- Pseudunio_flabellata **      
        +--+                             
           +-- Pseudunio_flabellatiformis **
           |                             
           +-- Pseudunio_homsensis **       

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

iTtTbiriSoTroTroSiD
3 3 1 3 138 8 2 2
  +----------- Cumberlandia          
--+                                  
  |  +-------- Margaritifera         
  +--+                               
     |  +----- Pseudunio_marocanus **   
     +--+                            
        |  +-- Pseudunio_auricularius **
        +--+                         
           +-- Pseudunio_homsensis **   

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 3 197 8 2 2
  +-------- Cumberlandia          
  |                               
--+-------- Margaritifera         
  |                               
  |  +----- Pseudunio_marocanus **   
  +--+                            
     |  +-- Pseudunio_auricularius **
     +--+                         
        +-- Pseudunio_homsensis **   

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

iTtTbiriSoTroTroSiD
3 3 1 3 628 8 2 2
  +----- Pseudunio_marocanus **   
--+                            
  |  +-- Pseudunio_auricularius **
  +--+                         
     +-- Pseudunio_homsensis **   

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
3 3 1 3 195 8 2 2
  +----------- Margaritifera         
--+                                  
  |  +-------- Cumberlandia          
  +--+                               
     |  +----- Pseudunio_marocanus **   
     +--+                            
        |  +-- Pseudunio_auricularius **
        +--+                         
           +-- Pseudunio_homsensis **   

Lopes-Lima et al., 2018 [Publ 3829], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 3 25 8 2 2
  +----------- Margaritifera         
--+                                  
  |  +-------- Cumberlandia          
  +--+                               
     |  +----- Pseudunio_marocanus **   
     +--+                            
        |  +-- Pseudunio_auricularius **
        +--+                         
           +-- Pseudunio_homsensis **   

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
3 3 1 3 197 8 2 2
  +----------- Cumberlandia          
--+                                  
  |  +-------- Margaritifera         
  +--+                               
     |  +----- Pseudunio_marocanus **   
     +--+                            
        |  +-- Pseudunio_auricularius **
        +--+                         
           +-- Pseudunio_homsensis **   

Lopes-Lima et al., 2018 [Publ 3829], fig. 3

iTtTbiriSoTroTroSiD
3 3 1 3 25 8 2 2
  +----------- Margaritifera         
--+                                  
  |  +-------- Cumberlandia          
  +--+                               
     |  +----- Pseudunio_marocanus **   
     +--+                            
        |  +-- Pseudunio_auricularius **
        +--+                         
           +-- Pseudunio_homsensis **   

Chen et al., 2025 [Publ 4455], fig. 1

iTtTbiriSoTroTroSiD
3 15 13 4 23 8 2 15
  +----------- Pseudunio_marocanus **      
  |                                     
  |        +-- Pseudunio_auricularius__1 **
--+        |                            
  |  +-----+-- Pseudunio_auricularius__2 **
  |  |     |                            
  +--+     +-- Pseudunio_homsensis **      
     |                                  
     |  +----- Gibbosula                
     +--+                               
        |  +-- Cumberlandia             
        +--+                            
           +-- Margaritifera            

Vikhrev et al., 2018 [Publ 3812], fig. 3

iTtTbiriSoTroTroSiD
3 3 1 3 10 7 2 2
  +----------- Margaritifera         
--+                                  
  |  +-------- Cumberlandia          
  +--+                               
     |  +----- Pseudunio_marocanus **   
     +--+                            
        |  +-- Pseudunio_auricularius **
        +--+                         
           +-- Pseudunio_homsensis **   

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iTtTbiriSoTroTroSiD
2 2 0 2 74 7 2 --
  +-- Pseudunio_auricularius **
--+                         
  +-- Pseudunio_marocanus **   

Sano et al., 2020 [Publ 4138], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 66 8 2 --
  +-- Pseudunio_auricularius **
--+                         
  +-- Pseudunio_marocanus **   

Wu et al., 2020 [Publ 4121], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 32 8 2 --
  +-- Pseudunio_auricularius **
--+                         
  +-- Pseudunio_marocanus **   

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
2 2 0 2 136 7 2 --
  +-- Pseudunio_auricularius **
--+                         
  +-- Pseudunio_marocanus **   

Wu et al., 2020 [Publ 4121], fig. S1

iTtTbiriSoTroTroSiD
2 2 0 2 32 8 2 --
  +-------- Margaritifera         
--+                               
  |  +----- Cumberlandia          
  +--+                            
     |  +-- Pseudunio_auricularius **
     +--+                         
        +-- Pseudunio_marocanus **   

Bolotov et al., 2015 [Publ 3646], fig. 2B

iTtTbiriSoTroTroSiD
2 2 0 2 7 6 2 --
  +-------- Cumberlandia          
--+                               
  |  +----- Margaritifera         
  +--+                            
     |  +-- Pseudunio_auricularius **
     +--+                         
        +-- Pseudunio_marocanus **   

Sano et al., 2020 [Publ 4138], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 66 8 2 --
  +-- Pseudunio_auricularius **
--+                         
  +-- Pseudunio_marocanus **   

Bolotov et al., 2016 [Publ 3743], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 28 7 2 --
  +-- Pseudunio_auricularius **
--+                         
  +-- Pseudunio_marocanus **   

Inoue et al., 2013 [Publ 3579], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 7 6 2 --
  +-------- Cumberlandia          
--+                               
  |  +----- Margaritifera         
  +--+                            
     |  +-- Pseudunio_auricularius **
     +--+                         
        +-- Pseudunio_marocanus **   

Guerra et al., 2019 [Publ 4168], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 39 4 2 --
  +-------- Cumberlandia          
--+                               
  |  +----- Margaritifera         
  +--+                            
     |  +-- Pseudunio_marocanus **   
     +--+                         
        +-- Pseudunio_auricularius **

Araujo et al., 2016 [Publ 3746], fig. 1_2b

iTtTbiriSoTroTroSiD
2 2 0 2 11 8 2 --
  +-------- Margaritifera         
--+                               
  |  +----- Cumberlandia          
  +--+                            
     |  +-- Pseudunio_marocanus **   
     +--+                         
        +-- Pseudunio_auricularius **

Araujo et al., 2009 [Publ 2395], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 6 5 2 --
  +-------- Cumberlandia          
--+                               
  |  +----- Margaritifera         
  +--+                            
     |  +-- Pseudunio_marocanus **   
     +--+                         
        +-- Pseudunio_auricularius **

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iTtTbiriSoTroTroSiD
2 2 0 2 74 7 2 --
  +----- Margaritifera         
--+                            
  |  +-- Pseudunio_auricularius **
  +--+                         
     +-- Pseudunio_marocanus **   

Huang et al., 2018 [Publ 3828], fig. 3_5

iTtTbiriSoTroTroSiD
2 2 0 2 27 8 2 --
  +-------- Margaritifera         
--+                               
  |  +----- Cumberlandia          
  +--+                            
     |  +-- Pseudunio_auricularius **
     +--+                         
        +-- Pseudunio_marocanus **   

Sano et al., 2020 [Publ 4138], fig. 4

iTtTbiriSoTroTroSiD
2 2 0 2 66 8 2 --
  +-- Pseudunio_auricularius **
--+                         
  +-- Pseudunio_marocanus **   

Takeuchi et al., 2015 [Publ 3661], fig. 2

iTtTbiriSoTroTroSiD
2 8 0 2 8 6 2 9
  +-------- Pseudunio_marocanus **   
--+                               
  |  +----- Pseudunio_auricularius **
  +--+                            
     |  +-- Cumberlandia          
     +--+                         
        +-- Margaritifera         

Bolotov et al., 2016 [Publ 3743], fig. 4

iTtTbiriSoTroTroSiD
2 2 0 2 28 7 2 --
  +-- Pseudunio_auricularius **
--+                         
  +-- Pseudunio_marocanus **   

Riccardi et al., 2020 [Publ 4089], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 17 0 0 --
  +-- Pseudunio_auricularius **
--+                         
  +-- Pseudunio_marocanus **   

Guerra et al., 2019 [Publ 4168], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 39 4 2 --
  +-------- Cumberlandia          
--+                               
  |  +----- Margaritifera         
  +--+                            
     |  +-- Pseudunio_marocanus **   
     +--+                         
        +-- Pseudunio_auricularius **

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
2 2 0 2 136 7 2 --
  +-- Pseudunio_auricularius **
--+                         
  +-- Pseudunio_marocanus **   

Araujo et al., 2016 [Publ 3746], fig. 2a

iTtTbiriSoTroTroSiD
2 2 0 2 9 8 2 --
  +-- Pseudunio_marocanus **   
--+                         
  +-- Pseudunio_auricularius **

Zieritz et al., 2021 [Publ 4231], fig. 3-M_4

iTtTbiriSoTroTroSiD
1 1 0 1 22 0 0 --
-- Pseudunio_marocanus **

Zieritz et al., 2021 [Publ 4218], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 30 1 1 --
  +-- Margaritifera         
--+                         
  +-- Pseudunio_auricularius **

Bolotov et al., 2015 [Publ 3646], fig. 2A

iTtTbiriSoTroTroSiD
1 1 0 1 8 7 2 --
  +----------- Margaritifera_margaritifera
  |                                       
  |     +----- Margaritifera_dahurica__1  
  |     |                                 
  +-----+----- Margaritifera_dahurica__2  
--+     |                                 
  |     |  +-- Pseudunio_auricularius **     
  |     +--+                              
  |        +-- Cumberlandia               
  |                                       
  |  +-------- Margaritifera_falcata      
  +--+                                    
     |  +----- Margaritifera_laevis       
     +--+                                 
        |  +-- Margaritifera_middendorffi 
        +--+                              
           +-- Margaritifera_marrianae    

Bashê, 2022 [Publ 4265], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 5 0 0 --
-- Pseudunio_homsensis **

Froufe et al., 2019 [Publ 3936], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 43 2 1 --
  +-- Pseudunio_marocanus **
--+                      
  +-- Margaritifera      

Walker et al., 2006 [Publ 2666], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 35 5 2 --
  +----- Cumberlandia          
--+                            
  |  +-- Pseudunio_auricularius **
  +--+                         
     +-- Margaritifera         

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iTtTbiriSoTroTroSiD
1 1 0 1 95 4 2 --
  +----- Cumberlandia       
--+                         
  |  +-- Pseudunio_marocanus **
  +--+                      
     +-- Margaritifera      

Huff et al., 2004 [Publ 2697], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 10 6 2 --
  +-- Pseudunio_auricularius **
  |                         
--+-- Cumberlandia          
  |                         
  +-- Margaritifera         

Huff et al., 2004 [Publ 2697], fig. 5-6

iTtTbiriSoTroTroSiD
1 1 0 1 11 6 2 --
  +----- Margaritifera         
--+                            
  |  +-- Pseudunio_auricularius **
  +--+                         
     +-- Cumberlandia          

Guerra et al., 2019 [Publ 4168], fig. 1-M

iTtTbiriSoTroTroSiD
1 1 0 1 21 2 2 --
  +----- Cumberlandia       
--+                         
  |  +-- Margaritifera      
  +--+                      
     +-- Pseudunio_marocanus **

Araujo et al., 2009 [Publ 2589], fig. 7C

iTtTbiriSoTroTroSiD
1 1 0 1 9 0 0 --
-- Pseudunio_auricularius **

Wu et al., 2021 [Publ 4366], fig. 3-BI

iTtTbiriSoTroTroSiD
1 1 0 1 73 4 2 --
  +----- Cumberlandia       
--+                         
  |  +-- Pseudunio_marocanus **
  +--+                      
     +-- Margaritifera      

Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1-M

iTtTbiriSoTroTroSiD
1 1 0 1 5 2 2 --
  +----- Margaritifera      
--+                         
  |  +-- Pseudunio_marocanus **
  +--+                      
     +-- Cumberlandia       

Prie et al., 2012 [Publ 3487], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 5 0 0 --
-- Pseudunio_auricularius **

Araujo et al., 2009 [Publ 2589], fig. 7B

iTtTbiriSoTroTroSiD
1 1 0 1 10 0 0 --
-- Pseudunio_auricularius **

Wu et al., 2022 [Publ 4238], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 73 4 2 --
  +----- Cumberlandia       
--+                         
  |  +-- Pseudunio_marocanus **
  +--+                      
     +-- Margaritifera      

Froufe et al., 2019 [Publ 3936], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 43 2 1 --
  +-- Pseudunio_marocanus **
--+                      
  +-- Margaritifera      

Machordom et al., 2003 [Publ 2688], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 3 2 2 --
  +----- Cumberlandia          
--+                            
  |  +-- Margaritifera         
  +--+                         
     +-- Pseudunio_auricularius **

Huang et al., 2019 [Publ 3843], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 55 4 2 --
  +----- Cumberlandia       
--+                         
  |  +-- Pseudunio_marocanus **
  +--+                      
     +-- Margaritifera      

Burzynski & Soroka, 2018 [Publ 3862], fig. 5a

iTtTbiriSoTroTroSiD
1 1 0 1 19 1 1 --
  +-- Pseudunio_marocanus **
--+                      
  +-- Cumberlandia       

Fukata & Iigo, 2020 [Publ 4106], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 53 1 1 --
  +-- Pseudunio_marocanus **
--+                      
  +-- Cumberlandia       

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iTtTbiriSoTroTroSiD
1 1 0 1 95 4 2 --
  +----- Cumberlandia       
--+                         
  |  +-- Pseudunio_marocanus **
  +--+                      
     +-- Margaritifera      

Zieritz et al., 2024 [Publ 4381], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 44 1 1 --
  +-- Margaritifera         
--+                         
  +-- Pseudunio_auricularius **

Wu et al., 2021 [Publ 4366], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 90 5 2 --
  +----- Margaritifera      
--+                         
  |  +-- Cumberlandia       
  +--+                      
     +-- Pseudunio_homsensis **

Cha et al., 2018 [Publ 3916], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 37 2 1 --
  +-- Pseudunio_marocanus **
--+                      
  +-- Margaritifera      

Machordom et al., 2003 [Publ 2688], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 2 1 1 --
  +-- Margaritifera         
--+                         
  +-- Pseudunio_auricularius **

Araujo et al., 2017 [Publ 3801], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 13 0 0 --
-- Pseudunio_marocanus **

Araujo et al., 2009 [Publ 2589], fig. 7A

iTtTbiriSoTroTroSiD
1 1 0 1 10 0 0 --
-- Pseudunio_auricularius **

Huff et al., 2004 [Publ 2697], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 7 5 2 --
  +-------- Margaritifera_margaritifera  
  |                                      
  |  +----- Cumberlandia                 
--+  |                                   
  |  |  +-- Pseudunio_auricularius **       
  |  |  |                                
  +--+  +-- Margaritifera_dahurica       
     |  |                                
     |  +-- Margaritifera_middendorffi__1
     |  |                                
     |  +-- Margaritifera_middendorffi__2
     +--+                                
        +-- Margaritifera_falcata__1     
        |                                
        +-- Margaritifera_falcata__2     
        |                                
        +-- Margaritifera_falcata__3     
        |                                
        +-- Margaritifera_falcata__4     

Guerra et al., 2019 [Publ 4168], fig. 2-M

iTtTbiriSoTroTroSiD
1 1 0 1 21 2 2 --
  +----- Pseudunio_marocanus **
--+                         
  |  +-- Margaritifera      
  +--+                      
     +-- Cumberlandia       

Huang et al., 2018 [Publ 3828], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 41 4 2 --
  +----- Cumberlandia       
--+                         
  |  +-- Pseudunio_marocanus **
  +--+                      
     +-- Margaritifera      

Wu et al., 2021 [Publ 4366], fig. 3-ML

iTtTbiriSoTroTroSiD
1 1 0 1 73 4 2 --
  +----- Cumberlandia       
--+                         
  |  +-- Pseudunio_marocanus **
  +--+                      
     +-- Margaritifera      

Lopes-Lima et al., 2017 [Publ 3792], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 31 2 1 --
  +-- Pseudunio_marocanus **
--+                      
  +-- Margaritifera      

Campbell & Lydeard, 2026 [Publ 4810], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 46 2 2 --
  +----- Pseudunio_marocanus **
--+                         
  |  +-- Cumberlandia       
  +--+                      
     +-- Margaritifera      

Froufe et al., 2019 [Publ 3936], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 18 0 0 --
-- Pseudunio_marocanus **

Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 9 4 2 --
  +----- Cumberlandia       
--+                         
  |  +-- Pseudunio_marocanus **
  +--+                      
     +-- Margaritifera      

Välilä et al., 2017 [Publ 3793], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 2 1 1 --
  +-- Margaritifera         
--+                         
  +-- Pseudunio_auricularius **

Froufe et al., 2019 [Publ 3936], fig. 3-M

iTtTbiriSoTroTroSiD
1 1 0 1 18 0 0 --
-- Pseudunio_marocanus **

Huff et al., 2004 [Publ 2697], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 11 6 2 --
  +-------- Margaritifera_dahurica        
  |                                       
  +-------- Margaritifera_margaritifera__1
  |                                       
  +-------- Margaritifera_margaritifera__2
  |                                       
--+-------- Margaritifera_margaritifera__3
  |                                       
  |     +-- Pseudunio_auricularius **        
  +-----+                                 
  |     +-- Cumberlandia                  
  |                                       
  |  +----- Margaritifera_falcata         
  +--+                                    
     |  +-- Margaritifera_middendorffi    
     +--+                                 
        +-- Margaritifera_marrianae       

Wu et al., 2021 [Publ 4366], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 68 4 2 --
  +----- Cumberlandia       
--+                         
  |  +-- Pseudunio_marocanus **
  +--+                      
     +-- Margaritifera      

Lopes-Lima et al., 2017 [Publ 3792], fig. 2-M

iTtTbiriSoTroTroSiD
1 1 0 1 9 0 0 --
-- Pseudunio_marocanus **

Dai et al., 2025 [Publ 4440], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 87 4 2 --
  +----- Cumberlandia       
--+                         
  |  +-- Pseudunio_marocanus **
  +--+                      
     +-- Margaritifera      

Takeuchi et al., 2015 [Publ 3661], fig. 3

iTtTbiriSoTroTroSiD
1 8 0 2 9 7 2 12
  +----- Pseudunio_auricularius__1 **
  |                               
--+----- Pseudunio_auricularius__2 **
  |                               
  |  +-- Cumberlandia             
  +--+                            
     +-- Margaritifera            

Lopes-Lima et al., 2017 [Publ 3734], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 16 1 1 --
  +-- Pseudunio_auricularius **
--+                         
  +-- Margaritifera         

Chen et al., 2025 [Publ 4448], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 85 4 2 --
  +----- Cumberlandia       
--+                         
  |  +-- Pseudunio_marocanus **
  +--+                      
     +-- Margaritifera      

Huang et al., 2019 [Publ 3843], fig. 3-M

iTtTbiriSoTroTroSiD
1 1 0 1 22 1 1 --
  +-- Pseudunio_marocanus **
--+                      
  +-- Margaritifera      

Prié & Puillandre, 2014 [Publ 3529], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 16 0 0 --
-- Pseudunio_auricularius **

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

iTtTbiriSoTroTroSiD
1 1 0 1 47 2 1 --
  +-- Pseudunio_marocanus **
--+                      
  +-- Margaritifera      

Wu et al., 2021 [Publ 4366], fig. 2_I

iTtTbiriSoTroTroSiD
1 1 0 1 90 5 2 --
  +----- Margaritifera      
--+                         
  |  +-- Pseudunio_homsensis **
  +--+                      
     +-- Cumberlandia       

Araujo et al., 2018 [Publ 3779], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 17 0 0 --
-- Pseudunio_auricularius **