MUSSELpdb
Taxonomy
Species & Genus Richness of
freshwater mussels
by Taxonomy
Species & Genus Richness of
freshwater bivalves
by Family
Geography
Species Richness & Endemism of
freshwater mussels
by Geography
Data last updated
Thu 19 March 2026
Publication Wu et al. (2023): cladograms
Complete Citation
Wu, R., L. Liu, L. Zhang, D. Jin, X. Wu et al. 2023. Molecular phylogeny and comparative morphology reveal the species validity and systematic position of Lepidodesma (Bivalvia: Unionidae). Organisms Diversity & Evolution (2023): 1-16. Available electronically.
Cladogram Figures
(click the figure number to see the cladogram)
Cladogram Figure 4_S5-aa
+-------------------------------------------------- Neotrigonia_margaritacea
|
| +----------------------------------------------- Echyridella_menziesii
| |
| | +-- Anodontites_trapesialis
--+ | +-----------------------------------------+
| | | +-- Mutela_dubia
| | |
| | | +-- Gibbosula_crassa
| | | +-----------+
+--+ | | +-- Gibbosula_rochechouartii
| | +--------------------------+
| | | | +----------- Cumberlandia_monodonta
| | | +--+
| | | | +-------- Pseudunio_marocanus
| | | +--+
| | | | +----- Margaritifera_falcata
| | | +--+
| | | | +-- Margaritifera_dahurica
+--+ | +--+
| | +-- Margaritifera_margaritifera
| |
| | +-- Quadrula_quadrula
| | +--------------------------+
| | | +-- Uniomerus_tetralasmus
| | |
| | +--------+ +-- Elliptio_complanata
| | | | +-----------------------+
| | | | | +-- Pleurobema_oviforme
| | | +--+
| | | | +----------------------- Amblema_plicata
| | | | |
| | | +--+ +-------------------- Popenaias_popeii
+--+ | | |
| | +--+ +----------------- Toxolasma_parvum
| | | |
| | | | +----- Potamilus_leptodon
| | +--+ +--------+
| | | | | +-- Potamilus_alatus
| | | | +--+
| | | | +-- Potamilus_streckersoni
| | +--+
| | | +----------- Leaunio_lienosus
| | | |
| | | | +----- Lampsilis_ornata
| | +--+ +--+
| | | | | +-- Lampsilis_cardium
| | | | +--+
| | +--+ +-- Lampsilis_cariosa
| | |
| | | +----- Venustaconcha_ellipsiformis
| | +--+
| | | +-- Lampsilis_powellii
| | +--+
| | +-- Lampsilis_siliquoidea
+--+
| +----- Chamberlainia_somsakpanhai
| +-----------------+
| | | +-- Sinohyriopsis_cumingii
| | +--+
| | +-- Sinohyriopsis_schlegelii
| |
| | +-- Hyriopsis_bialata
| | +--+
| +-----------+ | +-- Rectidens_sumatrensis
| | | +--------------+
| | | | | +-- Lens_contradens
| | | | +--+
| | | | +-- Physunio_superbus
| | | |
| | | | +----------- Potomida_littoralis
| | +--+ +-----+
| | | | | +-------- Pronodularia_japanensis
| | | | +--+
| | | | | +----- Lamprotula_leaii
| | | | +--+
| | | | | +-- Lamprotula_caveata
| | +--+ +--+
| | | +-- Lamprotula_cornuumlunae
| | |
| | | +-- Pseudodon_mekongi
| | | +-----------+
| | | | +-- Pilsbryoconcha_exilis
| | +--+
| | | +----------- Microcondylaea_bonellii
+--+ | |
| +--+ +-- Parvasolenaia_rivularis
| | +-----+
| | | +-- Ptychorhynchus_pfisteri
| +--+
| | +----- Sinosolenaia_carinata
| +--+
| | +-- Sinosolenaia_iridinea
| +--+
| +-- Sinosolenaia_oleivora
|
| +-------------------------------- Lamellidens_jenkinsianus
| |
| | +-- Lepidodesma_aligera
| | +--------------------------+
| | | +-- Lepidodesma_languilati
| | |
| | | +-------- Acuticosta_chinensis
| | | +--------+
| | | | | +----- Lanceolaria_gladiola
| | | | +--+
+--+ | | | +-- Lanceolaria_grayii
| | | +--+
| | | +-- Lanceolaria_lanceolata
| | |
| | | +-- Anemina_harlandi
| | | +--------+
| | +--------+ | +-- Cristaria_plicata
| | | | +--+
| | | | | | +-------- Beringiana_fukuharai
| | | | | +--+
+--+ | | | | +----- Sinanodonta_lucida
| | | | +--+
| | | | | +-- Sinanodonta_tumens
| | | | +--+
| | | | +-- Sinanodonta_woodiana
| | | |
| | +--+ +-- Brachyanodon_nuttalliana
| | | +-----+
| | | | +-- Pseudanodonta_complanata
| | | +--+
| | | | | +----- Anodonta_anatina
| | | | +--+
| | | | | +-- Anodonta_cygnea
| | | | +--+
| | | | +-- Anodonta_exulcerata
| | +--+
| | | +----- Pyganodon_grandis
| | | +--+
+--+ | | | +-- Utterbackia_peninsularis
| | | +--+
| | | +-- Utterbackia_imbecillis
| +--+
| | +-- Alasmidonta_varicosa
| | +--+
| | | +-- Platynaias_compressa
| +--+
| | +-- Prolasmidonta_heterodon
| +--+
| +-- Pterosyna_complanata
|
| +-------- Scriptolamprotula_tortuosa
| |
| +--------------+ +-- Scriptolamprotula_coreana
| | | +--+
| | | | +-- Scriptolamprotula_tientsinensis
| | +--+
| | | +-- Scriptolamprotula_polysticta
| | +--+
| | +-- Scriptolamprotula_scripta
| |
| | +-------------- Unio_tumidus
+--+ +-----+
| | | +----------- Unio_nanus
| | +--+
| | | +-------- Unio_elongatulus
| | +--+
| | | +----- Unio_pictorum
| | +--+
| | | +-- Unio_delphinus
| | +--+
+--+ +-- Unio_mancus
|
| +-- Nodularia_breviconcha
| +--+
| | +-- Nodularia_douglasiae
| +-----------+
| | | +-- Pseudocuneopsis_capitata
| | +--+
| | +-- Pseudocuneopsis_sichuanensis
+--+
| +-------------- Tchangsinaia_piscicula
| |
| | +----- Schistodesmus_spinosus
+--+ +-----+
| | | +-- Schistodesmus_lampreyanus
| | +--+
+--+ +-- Schistodesmus_xinyuensis
|
| +-------- Cuneopsis_rufescens
+--+
| +----- Cuneopsis_celtiformis
+--+
| +-- Cuneopsis_demangei
+--+
+-- Cuneopsis_heudei
Cladogram Figure 4_S5-nt
+----------------------------------------------------- Neotrigonia_margaritacea
|
| +-------------------------------------------------- Echyridella_menziesii
| |
| | +-- Anodontites_trapesialis
--+ | +--------------------------------------------+
| | | +-- Mutela_dubia
| | |
| | | +-- Gibbosula_crassa
| | | +-----------+
+--+ | | +-- Gibbosula_rochechouartii
| | +-----------------------------+
| | | | +----------- Cumberlandia_monodonta
| | | +--+
| | | | +-------- Pseudunio_marocanus
| | | +--+
| | | | +----- Margaritifera_falcata
| | | +--+
| | | | +-- Margaritifera_dahurica
+--+ | +--+
| | +-- Margaritifera_margaritifera
| |
| | +-- Quadrula_quadrula
| | +--------------------------+
| | | +-- Uniomerus_tetralasmus
| | |
| | +-----------+ +-- Elliptio_complanata
| | | | +-----------------------+
| | | | | +-- Pleurobema_oviforme
| | | +--+
| | | | +----------------------- Amblema_plicata
| | | | |
| | | +--+ +-------------------- Popenaias_popeii
+--+ | | |
| | +--+ +----------------- Toxolasma_parvum
| | | |
| | | | +----- Potamilus_leptodon
| | +--+ +--------+
| | | | | +-- Potamilus_alatus
| | | | +--+
| | | | +-- Potamilus_streckersoni
| | +--+
| | | +----------- Leaunio_lienosus
| | | |
| | | | +----- Lampsilis_ornata
| | +--+ +--+
| | | | | +-- Lampsilis_cardium
| | | | +--+
| | +--+ +-- Lampsilis_cariosa
| | |
| | | +----- Venustaconcha_ellipsiformis
| | +--+
| | | +-- Lampsilis_powellii
| | +--+
| | +-- Lampsilis_siliquoidea
+--+
| +----- Chamberlainia_somsakpanhai
| +-----------------+
| | | +-- Sinohyriopsis_cumingii
| | +--+
| | +-- Sinohyriopsis_schlegelii
| |
| | +-- Hyriopsis_bialata
| | +--+
| +--------------+ | +-- Rectidens_sumatrensis
| | | +--------------+
| | | | | +-- Lens_contradens
| | | | +--+
| | | | +-- Physunio_superbus
| | | |
| | | | +----------- Potomida_littoralis
| | +--+ +-----+
| | | | | +-------- Pronodularia_japanensis
| | | | +--+
| | | | | +----- Lamprotula_leaii
| | | | +--+
| | | | | +-- Lamprotula_caveata
| | +--+ +--+
| | | +-- Lamprotula_cornuumlunae
| | |
| | | +-- Pseudodon_mekongi
| | | +-----------+
| | | | +-- Pilsbryoconcha_exilis
| | | |
| | +--+ +-- Ptychorhynchus_pfisteri
+--+ | +--------+
| | | +-- Microcondylaea_bonellii
| +--+
| | +-------- Parvasolenaia_rivularis
| +--+
| | +----- Sinosolenaia_carinata
| +--+
| | +-- Sinosolenaia_iridinea
| +--+
| +-- Sinosolenaia_oleivora
|
| +----------------------------------- Lamellidens_jenkinsianus
| |
| | +-- Lepidodesma_aligera
| | +-----------------------------+
| | | +-- Lepidodesma_languilati
| | |
| | | +-------- Acuticosta_chinensis
| | | +--------------+
| | | | | +----- Lanceolaria_gladiola
| | | | +--+
+--+ | | | +-- Lanceolaria_grayii
| | | +--+
| | | +-- Lanceolaria_lanceolata
| | |
| | +-----+ +-------------- Cristaria_plicata
| | | | +-----+
| | | | | | +----------- Anemina_harlandi
| | | | | +--+
| | | | | | +-------- Beringiana_fukuharai
| | | | | +--+
+--+ | | | | +----- Sinanodonta_lucida
| | | | +--+
| | +--+ | +-- Sinanodonta_tumens
| | | +--+
| | | +-- Sinanodonta_woodiana
| | |
| | | +----------------- Brachyanodon_nuttalliana
| | | |
| | | +----------------- Pseudanodonta_complanata
| | | |
| | +--+ +-------------- Anodonta_cygnea
| | | |
| | | | +-- Anodonta_exulcerata
| | | +-----------+
| | | | +-- Anodonta_anatina
| | +--+
| | | +----- Pyganodon_grandis
+--+ | +-----+
| | | | +-- Utterbackia_imbecillis
| | | +--+
| +--+ +-- Utterbackia_peninsularis
| |
| | +-------- Pterosyna_complanata
| +--+
| | +----- Prolasmidonta_heterodon
| +--+
| | +-- Alasmidonta_varicosa
| +--+
| +-- Platynaias_compressa
|
| +-------- Scriptolamprotula_tortuosa
| |
| +-----------------+ +-- Scriptolamprotula_coreana
| | | +--+
| | | | +-- Scriptolamprotula_tientsinensis
| | +--+
| | | +-- Scriptolamprotula_polysticta
| | +--+
| | +-- Scriptolamprotula_scripta
| |
+--+ +----------- Unio_tumidus
| |
| +-----------+ +-------- Unio_nanus
| | | |
| | +--+ +----- Unio_elongatulus
| | | |
| | +--+ +-- Unio_delphinus
| | | |
| | +--+-- Unio_mancus
+--+ |
| +-- Unio_pictorum
|
| +-- Pseudocuneopsis_capitata
| +-----------------+
| | +-- Pseudocuneopsis_sichuanensis
| |
+--+ +-- Nodularia_breviconcha
| +--------------+
| | +-- Nodularia_douglasiae
| |
+--+ +----- Tchangsinaia_piscicula
| +--------+
| | | +-- Schistodesmus_xinyuensis
| | +--+
+--+ +-- Schistodesmus_spinosus
|
| +----------- Schistodesmus_lampreyanus
+--+
| +-------- Cuneopsis_rufescens
+--+
| +----- Cuneopsis_celtiformis
+--+
| +-- Cuneopsis_demangei
+--+
+-- Cuneopsis_heudei
Cladogram Figure 5
+-- Margaritifera_margaritifera
+-----------------+
| +-- Margaritifera_dahurica
|
| +-- Elliptio_complanata
| +--------+
| | +-- Quadrula_quadrula
--+ +-----+
| | | +-------- Obliquaria_reflexa
| | +--+
| | | +----- Truncilla_truncata
| | +--+
| | | +-- Lampsilis_cardium
| | +--+
+--+ +-- Potamilus_alatus
|
| +-- Lepidodesma_languilati
| +-----------+
| | +-- Lepidodesma_aligera
| |
| | +-------- Anemina_harlandi
+--+ +--+
| | | +----- Cristaria_plicata
| | +--+
| | | +-- Sinanodonta_tumens
| | +--+
+--+ +-- Sinanodonta_woodiana
|
| +-- Platynaias
| +-----+
| | +-- Platynaias_compressa
+--+
| +----- Anodonta_anatina
+--+
| +-- Pseudanodonta_complanata
+--+
+-- Anodonta_cygnea
