Bivalvia | Unionida | Unionoidea | Unionidae | Ambleminae | Popenaiadini

genus Popenaias Frierson, 1927: cladomics

4 Recent valid species (4 total species)

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 4 (4 total)
analyzed in 45 cladograms in 28 publications
maximum number of terminals in an analysis = 3 (mean = 1.13, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 3
maximum number of included terminals = 3, (mean = 3 median = 3)
proportion of cladograms that support basic monophyly = 1

Cladograms

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
3 3 1 3 628 3 2 0 
  +----- Popenaias_popeii **  
--+                        
  |  +-- Popenaias_berezai ** 
  +--+                     
     +-- Popenaias_semirasa **

Inoue et al., 2020 [Publ 4128], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 3 20 2 1 0 
  +-------- Psoronaias        
--+                           
  |  +----- Popenaias_popeii **  
  +--+                        
     |  +-- Popenaias_berezai ** 
     +--+                     
        +-- Popenaias_semirasa **

Inoue et al., 2020 [Publ 4128], fig. 3

iTtTbiriSoTroTroSiD
3 3 1 3 19 2 1 0 
  +-------- Psoronaias        
--+                           
  |  +----- Popenaias_popeii **  
  +--+                        
     |  +-- Popenaias_berezai ** 
     +--+                     
        +-- Popenaias_semirasa **

Zanatta & Murphy, 2006 [Publ 2672], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 56 0 0 -- 
-- Popenaias_popeii **

Freeman et al., 2025 [Publ 4443], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 11 1 1 -- 
  +-- Nephronaias     
--+                   
  +-- Popenaias_popeii **

Campbell & Lydeard, 2012 [Publ 3308], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 41 0 0 -- 
-- Popenaias_popeii **

Inoue et al., 2018 [Publ 3858], fig. 1_3a-c

iTtTbiriSoTroTroSiD
1 1 0 1 104 0 0 -- 
-- Popenaias_popeii **

Dai et al., 2025 [Publ 4440], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 87 0 0 -- 
-- Popenaias_popeii **

Burlakova et al., 2012 [Publ 3486], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 40 0 0 -- 
-- Popenaias_popeii **

Campbell & Lydeard, 2012 [Publ 3308], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 41 0 0 -- 
-- Popenaias_popeii **

Wu et al., 2021 [Publ 4366], fig. 3-ML

iTtTbiriSoTroTroSiD
1 1 0 1 73 0 0 -- 
-- Popenaias_popeii **

Chapman et al., 2008 [Publ 2667], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 20 0 0 -- 
-- Popenaias_popeii **

Chapman et al., 2008 [Publ 2744], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 20 0 0 -- 
-- Popenaias_popeii **

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

iTtTbiriSoTroTroSiD
1 1 0 1 140 0 0 -- 
-- Popenaias_popeii **

Smith et al., 2020 [Publ 4129], fig. 2-3

iTtTbiriSoTroTroSiD
1 1 0 1 27 1 1 -- 
  +-- Popenaias_popeii **
--+                   
  +-- Psoronaias

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iTtTbiriSoTroTroSiD
1 1 0 1 95 0 0 -- 
-- Popenaias_popeii **

Freeman et al., 2025 [Publ 4443], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 5 3 2 -- 
  +----- Popenaias_popeii **
--+                      
  |  +-- Nephronaias     
  +--+                   
     +-- Psoronaias

Campbell et al., 2005 [Publ 2660], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 101 0 0 -- 
-- Popenaias_popeii **

Burlakova et al., 2019 [Publ 3899], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 32 0 0 -- 
-- Popenaias_popeii **

Pfeiffer et al., 2019 [Publ 3866], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 30 2 1 -- 
  +-- Popenaias_popeii **
--+                   
  +-- Psoronaias

Campbell et al., 2005 [Publ 2660], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 101 0 0 -- 
-- Popenaias_popeii **

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iTtTbiriSoTroTroSiD
1 1 0 1 95 0 0 -- 
-- Popenaias_popeii **

Chen et al., 2025 [Publ 4448], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 85 0 0 -- 
-- Popenaias_popeii **

Campbell & Lydeard, 2012 [Publ 3308], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 42 0 0 -- 
-- Popenaias_popeii **

Dai et al., 2024 [Publ 4358], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 52 0 0 -- 
-- Popenaias_popeii **

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iTtTbiriSoTroTroSiD
1 1 0 1 91 0 0 -- 
-- Popenaias_popeii **

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 56 1 1 -- 
  +-- Psoronaias      
--+                   
  +-- Popenaias_popeii **

Dai et al., 2023 [Publ 4285], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 52 0 0 -- 
-- Popenaias_popeii **

Wu et al., 2021 [Publ 4366], fig. 3-BI

iTtTbiriSoTroTroSiD
1 1 0 1 73 0 0 -- 
-- Popenaias_popeii **

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iTtTbiriSoTroTroSiD
1 1 0 1 91 0 0 -- 
-- Popenaias_popeii **

Mejia-Trujillo & Smith, 2022 [Publ 4274], fig. 2-F

iTtTbiriSoTroTroSiD
1 1 0 1 14 0 0 -- 
-- Popenaias_popeii **

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iTtTbiriSoTroTroSiD
1 1 0 1 56 1 1 -- 
  +-- Psoronaias      
--+                   
  +-- Popenaias_popeii **

Campbell & Lydeard, 2012 [Publ 3309], fig. 1-2

iTtTbiriSoTroTroSiD
1 1 0 1 72 0 0 -- 
-- Popenaias_popeii **

Campbell & Lydeard, 2012 [Publ 3308], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 41 0 0 -- 
-- Popenaias_popeii **

Wu et al., 2024 [Publ 4388], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 52 0 0 -- 
-- Popenaias_popeii **

Freeman et al., 2025 [Publ 4443], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 10 1 1 -- 
  +-- Nephronaias     
--+                   
  +-- Popenaias_popeii **

Freeman et al., 2025 [Publ 4443], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 11 1 1 -- 
  +-- Nephronaias     
--+                   
  +-- Popenaias_popeii **

Wu et al., 2021 [Publ 4366], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 68 0 0 -- 
-- Popenaias_popeii **

Wu et al., 2022 [Publ 4238], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 73 0 0 -- 
-- Popenaias_popeii **

Smith et al., 2019 [Publ 3951], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 40 0 0 -- 
-- Popenaias_popeii **

Smith et al., 2020 [Publ 4129], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 27 1 1 -- 
  +-- Popenaias_popeii **
--+                   
  +-- Psoronaias

Neemuchwala et al., 2023 [Publ 4292], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 26 0 0 -- 
-- Popenaias_popeii **

Zanatta & Murphy, 2006 [Publ 2672], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 55 0 0 -- 
-- Popenaias_popeii **

Campbell & Lydeard, 2012 [Publ 3309], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 72 0 0 -- 
-- Popenaias_popeii **

Teiga-Teixeira et al., 2020 [Publ 4173], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 18 0 0 -- 
-- Popenaias_popeii **