Bivalvia | Unionida | Unionoidea | Unionidae | Gonideinae | Lamprotulini

genus Lamprotula Simpson, 1900: cladomics

17 Recent valid species (26 total species)

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 17 (26 total)
analyzed in 146 cladograms in 91 publications
maximum number of terminals in an analysis = 4 (mean = 1.67, median = 2)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 74
maximum number of included terminals = 4, (mean = 2.32 median = 2)
proportion of cladograms that support basic monophyly = 0.93


Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Lamprotula cladogram distances
PCoA Ordination

NJ Tree of Lamprotula cladogram distances
NJ Tree


Cladograms

Huang et al., 2019 [Publ 3843], fig. S1

** Top Monophyly Cladogram **

iTtTbiriSoTroTroSiD
3 3 2 6 44 0 0 0
  +-------- Lamprotula_leaii **          
  |                                   
--+  +----- Lamprotula_cornuumlunae__1 **
  |  |                                
  |  +----- Lamprotula_cornuumlunae__3 **
  +--+                                
     +----- Lamprotula_cornuumlunae__4 **
     |                                
     |  +-- Lamprotula_cornuumlunae__2 **
     +--+                             
        +-- Lamprotula_caveata **        

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
4 4 3 5 627 4 2 4
           +-- Pronodularia           
  +--------+                          
  |        +-- Potomida               
  |                                   
--+        +-- Lamprotula_caveata **     
  |  +-----+                          
  |  |     +-- Lamprotula_cornuumlunae **
  +--+                                
     |  +----- Lamprotula_leaii__2 **    
     +--+                             
        |  +-- Lamprotula_leaii__1 **    
        +--+                          
           +-- Lamprotula_seomjinensis **

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

iTtTbiriSoTroTroSiD
3 3 2 4 138 3 1 4
  +----------- Potomida               
--+                                   
  |  +-------- Lamprotula_leaii **       
  +--+                                
     |  +----- Lamprotula_caveata__2 **  
     +--+                             
        |  +-- Lamprotula_cornuumlunae **
        +--+                          
           +-- Lamprotula_caveata__1 **  

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iTtTbiriSoTroTroSiD
3 3 1 3 93 2 2 3
  +----------- Potomida               
--+                                   
  |  +-------- Pronodularia           
  +--+                                
     |  +----- Lamprotula_leaii **       
     +--+                             
        |  +-- Lamprotula_caveata **     
        +--+                          
           +-- Lamprotula_cornuumlunae **

Zhang et al., 2024 [Publ 4397], fig. 5

iTtTbiriSoTroTroSiD
3 3 1 3 24 2 2 3
  +----------- Potomida               
--+                                   
  |  +-------- Pronodularia           
  +--+                                
     |  +----- Lamprotula_leaii **       
     +--+                             
        |  +-- Lamprotula_caveata **     
        +--+                          
           +-- Lamprotula_cornuumlunae **

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iTtTbiriSoTroTroSiD
3 3 1 3 89 2 2 3
  +----------- Potomida               
--+                                   
  |  +-------- Pronodularia           
  +--+                                
     |  +----- Lamprotula_leaii **       
     +--+                             
        |  +-- Lamprotula_caveata **     
        +--+                          
           +-- Lamprotula_cornuumlunae **

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 3 197 2 2 3
        +-- Potomida               
  +-----+                          
  |     +-- Pronodularia           
--+                                
  |  +----- Lamprotula_leaii **       
  +--+                             
     |  +-- Lamprotula_caveata **     
     +--+                          
        +-- Lamprotula_cornuumlunae **

Wu et al., 2026 [Publ 4811], fig. 4

iTtTbiriSoTroTroSiD
3 3 1 3 92 3 2 3
  +----------- Potomida               
--+                                   
  |  +-------- Pronodularia           
  +--+                                
     |  +----- Lamprotula_leaii **       
     +--+                             
        |  +-- Lamprotula_caveata **     
        +--+                          
           +-- Lamprotula_cornuumlunae **

Wu et al., 2020 [Publ 4147], fig. 3

iTtTbiriSoTroTroSiD
3 3 2 4 27 1 1 4
  +----------- Pronodularia           
--+                                   
  |  +-------- Lamprotula_leaii **       
  +--+                                
     |  +----- Lamprotula_caveata__1 **  
     +--+                             
        |  +-- Lamprotula_cornuumlunae **
        +--+                          
           +-- Lamprotula_caveata__2 **  

Zhou et al., 2007 [Publ 2658], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 3 15 0 0 3
  +----- Lamprotula_leaii **       
--+                             
  |  +-- Lamprotula_caveata **     
  +--+                          
     +-- Lamprotula_cornuumlunae **

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
3 3 1 3 195 2 2 3
        +-- Potomida               
  +-----+                          
  |     +-- Pronodularia           
--+                                
  |  +----- Lamprotula_leaii **       
  +--+                             
     |  +-- Lamprotula_caveata **     
     +--+                          
        +-- Lamprotula_cornuumlunae **

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
3 3 2 5 119 2 2 4
  +-------------- Pronodularia           
  |                                      
--+  +----------- Potomida               
  |  |                                   
  +--+  +-------- Lamprotula_caveata **     
     |  |                                
     +--+  +----- Lamprotula_leaii__2 **    
        |  |                             
        +--+----- Lamprotula_leaii__3 **    
           |                             
           |  +-- Lamprotula_seomjinensis **
           +--+                          
              +-- Lamprotula_leaii__1 **    

Bolotov et al., 2020 [Publ 4100], fig. S1

iTtTbiriSoTroTroSiD
3 5 0 3 108 2 2 5
  +-- Lamprotula_leaii **  
  |                     
  +-- Lamprotula **        
  |                     
--+-- Lamprotula_caveata **
  |                     
  +-- Potomida          
  |                     
  +-- Pronodularia      

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iTtTbiriSoTroTroSiD
3 3 1 3 93 2 2 3
  +----------- Potomida               
--+                                   
  |  +-------- Pronodularia           
  +--+                                
     |  +----- Lamprotula_leaii **       
     +--+                             
        |  +-- Lamprotula_caveata **     
        +--+                          
           +-- Lamprotula_cornuumlunae **

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
3 3 1 3 197 2 2 3
        +-- Potomida               
  +-----+                          
  |     +-- Pronodularia           
--+                                
  |  +----- Lamprotula_leaii **       
  +--+                             
     |  +-- Lamprotula_caveata **     
     +--+                          
        +-- Lamprotula_cornuumlunae **

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iTtTbiriSoTroTroSiD
3 3 1 3 89 2 2 3
  +----------- Potomida               
--+                                   
  |  +-------- Pronodularia           
  +--+                                
     |  +----- Lamprotula_leaii **       
     +--+                             
        |  +-- Lamprotula_caveata **     
        +--+                          
           +-- Lamprotula_cornuumlunae **

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

iTtTbiriSoTroTroSiD
3 3 2 5 28 2 2 3
  +-------------- Pronodularia           
  |                                      
--+  +----------- Potomida               
  |  |                                   
  +--+  +-------- Lamprotula_leaii **       
     |  |                                
     +--+  +----- Lamprotula_caveata__3 **  
        |  |                             
        +--+  +-- Lamprotula_cornuumlunae **
           |  |                          
           +--+-- Lamprotula_caveata__1 **  
              |                          
              +-- Lamprotula_caveata__2 **  

Wu et al., 2024 [Publ 4388], fig. 6

iTtTbiriSoTroTroSiD
3 3 1 3 50 2 2 3
  +----------- Potomida               
--+                                   
  |  +-------- Pronodularia           
  +--+                                
     |  +----- Lamprotula_leaii **       
     +--+                             
        |  +-- Lamprotula_caveata **     
        +--+                          
           +-- Lamprotula_cornuumlunae **

Lopes-Lima et al., 2020 [Publ 4080], fig. 16

iTtTbiriSoTroTroSiD
3 3 2 4 5 1 1 5
  +----------- Pronodularia           
--+                                   
  |  +-------- Lamprotula_caveata **     
  +--+                                
     |  +----- Lamprotula_leaii__2 **    
     +--+                             
        |  +-- Lamprotula_seomjinensis **
        +--+                          
           +-- Lamprotula_leaii__1 **    

Bolotov et al., 2020 [Publ 4100], fig. 1

iTtTbiriSoTroTroSiD
3 16 0 3 114 2 2 16
  +-- Sinosolenaia_iridinea    
  |                            
  +-- Sinosolenaia_recognita__1
  |                            
  +-- Sinosolenaia_carinata    
  |                            
  +-- Leguminaia               
  |                            
  +-- Microcondylaea           
  |                            
  +-- Gonidea                  
  |                            
  +-- Ptychorhynchus           
  |                            
  +-- Parvasolenaia            
  |                            
--+-- Lamprotula_leaii **         
  |                            
  +-- Lamprotula **               
  |                            
  +-- Lamprotula_caveata **       
  |                            
  +-- Potomida                 
  |                            
  +-- Pronodularia             
  |                            
  +-- Sinohyriopsis            
  |                            
  +-- Chamberlainia            
  |                            
  +-- Sinosolenaia_recognita__2
  |                            
  +-- Sinosolenaia_oleivora    

Zhou et al., 2007 [Publ 2658], fig. 1

iTtTbiriSoTroTroSiD
3 4 2 3 18 0 0 4
  +-------- Lamprotula_leaii **       
--+                                
  |  +----- Lamprotula_caveata **     
  +--+                             
     |  +-- Gibbosula              
     +--+                          
        +-- Lamprotula_cornuumlunae **

Wu et al., 2020 [Publ 4147], fig. 4

iTtTbiriSoTroTroSiD
3 3 1 3 11 1 1 3
  +-------- Potomida               
--+                                
  |  +----- Lamprotula_leaii **       
  +--+                             
     |  +-- Lamprotula_caveata **     
     +--+                          
        +-- Lamprotula_cornuumlunae **

Huang et al., 2019 [Publ 3843], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 4 42 0 0 4
  +----- Lamprotula_leaii **       
  |                             
--+  +-- Lamprotula_caveata__1 **  
  |  |                          
  +--+-- Lamprotula_cornuumlunae **
     |                          
     +-- Lamprotula_caveata__2 **  

Froufe et al., 2019 [Publ 3936], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 42 2 2 --
  +-------- Potomida          
--+                           
  |  +----- Pronodularia      
  +--+                        
     |  +-- Lamprotula_leaii **  
     +--+                     
        +-- Lamprotula_caveata **

Dai et al., 2025 [Publ 4441], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 45 2 2 --
  +-------- Potomida          
--+                           
  |  +----- Pronodularia      
  +--+                        
     |  +-- Lamprotula_caveata **
     +--+                     
        +-- Lamprotula_leaii **  

Wu et al., 2018 [Publ 3836], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 51 1 1 --
  +----- Potomida          
--+                        
  |  +-- Lamprotula_caveata **
  +--+                     
     +-- Lamprotula_leaii **  

Zhang et al., 2024 [Publ 4397], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 6 0 0 --
  +-- Lamprotula_caveata **
--+                     
  +-- Lamprotula_leaii **  

Dai et al., 2024 [Publ 4389], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 44 2 2 --
     +-- Lamprotula_caveata **
  +--+                     
  |  +-- Lamprotula_leaii **  
--+                        
  |  +-- Pronodularia      
  +--+                     
     +-- Potomida          

Wu et al., 2021 [Publ 4215], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 22 2 2 --
     +-- Lamprotula_caveata **
  +--+                     
  |  +-- Lamprotula_leaii **  
--+                        
  |  +-- Potomida          
  +--+                     
     +-- Pronodularia      

Konopleva et al., 2023 [Publ 4269], fig. S1_4

iTtTbiriSoTroTroSiD
2 2 0 2 54 2 2 --
  +-------- Potomida          
--+                           
  |  +----- Pronodularia      
  +--+                        
     |  +-- Lamprotula_caveata **
     +--+                     
        +-- Lamprotula_leaii **  

Wu et al., 2021 [Publ 4366], fig. 3-BI

iTtTbiriSoTroTroSiD
2 2 0 2 72 2 2 --
  +-------- Potomida          
--+                           
  |  +----- Pronodularia      
  +--+                        
     |  +-- Lamprotula_caveata **
     +--+                     
        +-- Lamprotula_leaii **  

Wu et al., 2020 [Publ 4121], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 32 1 1 --
  +----- Potomida          
--+                        
  |  +-- Lamprotula_caveata **
  +--+                     
     +-- Lamprotula_leaii **  

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
2 2 0 2 136 2 2 --
     +-- Lamprotula_caveata **
  +--+                     
  |  +-- Lamprotula_leaii **  
--+                        
  |  +-- Potomida          
  +--+                     
     +-- Pronodularia      

Wu et al., 2022 [Publ 4238], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 72 2 2 --
  +-------- Potomida          
--+                           
  |  +----- Pronodularia      
  +--+                        
     |  +-- Lamprotula_caveata **
     +--+                     
        +-- Lamprotula_leaii **  

Wu et al., 2020 [Publ 4121], fig. S1

iTtTbiriSoTroTroSiD
2 2 0 2 32 1 1 --
  +----- Potomida          
--+                        
  |  +-- Lamprotula_caveata **
  +--+                     
     +-- Lamprotula_leaii **  

Matos et al., 2024 [Publ 4383], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 18 3 2 --
  +-------- Potomida          
--+                           
  |  +----- Pronodularia      
  +--+                        
     |  +-- Lamprotula_leaii **  
     +--+                     
        +-- Lamprotula_caveata **

Bolotov et al., 2017 [Publ 3764], fig. S1

iTtTbiriSoTroTroSiD
2 13 0 2 122 2 2 12
  +-- Chamberlainia         
  |                         
  +-- Sinohyriopsis         
  |                         
  +-- Lamprotula_caveata **    
  |                         
  +-- Lamprotula_leaii **      
  |                         
  +-- Potomida              
  |                         
  +-- Pronodularia          
  |                         
--+-- Gonidea               
  |                         
  +-- Leguminaia_wheatleyi  
  |                         
  +-- Leguminaia            
  |                         
  +-- Microcondylaea        
  |                         
  +-- Sinosolenaia_carinata 
  |                         
  +-- Sinosolenaia_oleivora 
  |                         
  +-- Sinosolenaia_recognita

Froufe et al., 2019 [Publ 3936], fig. 6

iTtTbiriSoTroTroSiD
2 2 0 2 42 2 2 --
  +-------- Potomida          
--+                           
  |  +----- Pronodularia      
  +--+                        
     |  +-- Lamprotula_leaii **  
     +--+                     
        +-- Lamprotula_caveata **

Huang et al., 2019 [Publ 3843], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 54 2 2 --
  +-------- Potomida          
--+                           
  |  +----- Pronodularia      
  +--+                        
     |  +-- Lamprotula_caveata **
     +--+                     
        +-- Lamprotula_leaii **  

Chen et al., 2025 [Publ 4663], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 46 2 2 --
  +-------- Potomida          
--+                           
  |  +----- Pronodularia      
  +--+                        
     |  +-- Lamprotula_caveata **
     +--+                     
        +-- Lamprotula_leaii **  

Fukata & Iigo, 2020 [Publ 4106], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 52 2 2 --
     +-- Lamprotula_caveata **
  +--+                     
  |  +-- Lamprotula_leaii **  
--+                        
  |  +-- Pronodularia      
  +--+                     
     +-- Potomida          

Liu et al., 2024 [Publ 4380], fig. 5

iTtTbiriSoTroTroSiD
2 2 0 2 34 1 1 --
  +----- Potomida          
--+                        
  |  +-- Lamprotula_caveata **
  +--+                     
     +-- Lamprotula_leaii **  

Lopes-Lima et al., 2020 [Publ 4080], fig. 14

iTtTbiriSoTroTroSiD
2 2 0 2 4 0 0 --
  +-- Lamprotula_leaii **       
--+                          
  +-- Lamprotula_seomjinensis **

Dai et al., 2023 [Publ 4285], fig. 3-BI

iTtTbiriSoTroTroSiD
2 2 0 2 43 2 2 --
  +-------- Potomida          
--+                           
  |  +----- Pronodularia      
  +--+                        
     |  +-- Lamprotula_leaii **  
     +--+                     
        +-- Lamprotula_caveata **

Liu et al., 2024 [Publ 4380], fig. 4

iTtTbiriSoTroTroSiD
2 2 0 2 10 0 0 --
  +-- Lamprotula_caveata **
--+                     
  +-- Lamprotula_leaii **  

Wu et al., 2021 [Publ 4366], fig. 6

iTtTbiriSoTroTroSiD
2 2 0 2 89 2 2 --
  +-------- Pronodularia      
--+                           
  |  +----- Potomida          
  +--+                        
     |  +-- Lamprotula_leaii **  
     +--+                     
        +-- Lamprotula_caveata **

Cha et al., 2018 [Publ 3916], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 36 2 2 --
  +-------- Potomida          
--+                           
  |  +----- Pronodularia      
  +--+                        
     |  +-- Lamprotula_caveata **
     +--+                     
        +-- Lamprotula_leaii **  

Wang et al., 2018 [Publ 3865], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 29 1 1 --
  +----- Potomida          
--+                        
  |  +-- Lamprotula_caveata **
  +--+                     
     +-- Lamprotula_leaii **  

Dai et al., 2023 [Publ 4285], fig. 3-ML

iTtTbiriSoTroTroSiD
2 2 0 2 43 2 2 --
  +-------- Potomida          
--+                           
  |  +----- Pronodularia      
  +--+                        
     |  +-- Lamprotula_leaii **  
     +--+                     
        +-- Lamprotula_caveata **

Huang et al., 2018 [Publ 3828], fig. 6

iTtTbiriSoTroTroSiD
2 2 0 2 40 2 2 --
  +-------- Potomida          
--+                           
  |  +----- Pronodularia      
  +--+                        
     |  +-- Lamprotula_caveata **
     +--+                     
        +-- Lamprotula_leaii **  

Wu et al., 2021 [Publ 4366], fig. 3-ML

iTtTbiriSoTroTroSiD
2 2 0 2 72 2 2 --
  +-------- Potomida          
--+                           
  |  +----- Pronodularia      
  +--+                        
     |  +-- Lamprotula_caveata **
     +--+                     
        +-- Lamprotula_leaii **  

Campbell & Lydeard, 2026 [Publ 4810], fig. 4

iTtTbiriSoTroTroSiD
2 2 0 2 45 1 1 --
  +----- Potomida          
--+                        
  |  +-- Lamprotula_caveata **
  +--+                     
     +-- Lamprotula_leaii **  

Chen et al., 2020 [Publ 4170], fig. 1

iTtTbiriSoTroTroSiD
2 34 0 2 32 0 0 33
                       +----- Prolasmidonta           
                    +--+                              
                    |  |  +-- Alasmidonta             
                    |  +--+                           
                    |     +-- Anemina                 
  +-----------------+                                 
  |                 |     +-- Lamprotula_caveata **      
  |                 |  +--+                           
  |                 |  |  +-- Chamberlainia           
  |                 +--+                              
  |                    |  +-- Pseudodon               
  |                    +--+                           
  |                       +-- Pilsbryoconcha          
  |                                                   
  |                    +----- Schistodesmus           
  |              +-----+                              
  |              |     |  +-- Lepidodesma             
--+              |     +--+                           
  |              |        +-- Scriptolamprotula       
  |  +-----------+                                    
  |  |           |     +----- Platynaias              
  |  |           |  +--+                              
  |  |           |  |  |  +-- Utterbackia_imbecillis  
  |  |           |  |  +--+                           
  |  |           +--+     +-- Pyganodon               
  |  |              |                                 
  |  |              |  +----- Sinanodonta             
  |  |              +--+                              
  |  |                 |  +-- Cristaria               
  +--+                 +--+                           
     |                    +-- Anodonta                
     |                                                
     |                    +-- Utterbackia_peninsularis
     |           +--------+                           
     |           |        +-- Potamilus_alatus        
     |  +--------+                                    
     |  |        |  +-------- Lamprotula_leaii **        
     |  |        +--+                                 
     |  |           |  +----- Sinohyriopsis           
     |  |           +--+                              
     +--+              |  +-- Parvasolenaia           
        |              +--+                           
        |                 +-- Sinosolenaia            
        |                                             
        |  +----------------- Gibbosula               
        +--+                                          
           |  +-------------- Quadrula                
           +--+                                       
              |  +----------- Toxolasma               
              +--+                                    
                 |  +-------- Lampsilis_ornata        
                 +--+                                 
                    |  +----- Potamilus_leptodon      
                    +--+                              
                       |  +-- Lampsilis_siliquoidea   
                       +--+                           
                          +-- Lampsilis_powellii      

Matos et al., 2024 [Publ 4384], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 18 3 2 --
  +-------- Potomida          
--+                           
  |  +----- Pronodularia      
  +--+                        
     |  +-- Lamprotula_leaii **  
     +--+                     
        +-- Lamprotula_caveata **

Wu et al., 2024 [Publ 4350], fig. 5a

iTtTbiriSoTroTroSiD
2 2 0 2 34 1 1 --
  +----- Potomida          
--+                        
  |  +-- Lamprotula_caveata **
  +--+                     
     +-- Lamprotula_leaii **  

Dai et al., 2023 [Publ 4285], fig. 5

iTtTbiriSoTroTroSiD
2 2 0 2 51 2 2 --
  +-------- Potomida          
--+                           
  |  +----- Pronodularia      
  +--+                        
     |  +-- Lamprotula_caveata **
     +--+                     
        +-- Lamprotula_leaii **  

Ouyang et al., 2015 [Publ 4493], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 29 0 0 --
  +-- Lamprotula_caveata **
--+                     
  +-- Lamprotula_leaii **  

Dai et al., 2024 [Publ 4386], fig. 6

iTtTbiriSoTroTroSiD
2 2 0 2 40 2 2 --
  +-------- Potomida          
--+                           
  |  +----- Pronodularia      
  +--+                        
     |  +-- Lamprotula_caveata **
     +--+                     
        +-- Lamprotula_leaii **  

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iTtTbiriSoTroTroSiD
2 2 0 2 78 2 2 --
     +-- Lamprotula_caveata **
  +--+                     
  |  +-- Lamprotula_leaii **  
--+                        
  |  +-- Potomida          
  +--+                     
     +-- Pronodularia      

Hou et al., 2024 [Publ 4400], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 13 0 0 --
  +-- Lamprotula_caveata **
--+                     
  +-- Lamprotula_leaii **  

Wu et al., 2021 [Publ 4366], fig. 4

iTtTbiriSoTroTroSiD
2 2 0 2 67 1 1 --
  +----- Potomida          
--+                        
  |  +-- Lamprotula_caveata **
  +--+                     
     +-- Lamprotula_leaii **  

Wu et al., 2024 [Publ 4350], fig. 5c

iTtTbiriSoTroTroSiD
2 2 0 2 34 1 1 --
  +-- Lamprotula_caveata **
--+                     
  +-- Lamprotula_leaii **  

Zhou et al., 2017 [Publ 3788], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 14 0 0 --
  +-- Lamprotula_caveata **
--+                     
  +-- Lamprotula_leaii **  

Dai et al., 2025 [Publ 4440], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 86 2 2 --
  +-------- Potomida          
--+                           
  |  +----- Pronodularia      
  +--+                        
     |  +-- Lamprotula_caveata **
     +--+                     
        +-- Lamprotula_leaii **  

Wu et al., 2024 [Publ 4379], fig. 4_6

iTtTbiriSoTroTroSiD
2 2 0 2 21 1 1 --
  +----- Potomida          
--+                        
  |  +-- Lamprotula_caveata **
  +--+                     
     +-- Lamprotula_leaii **  

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
2 2 0 2 136 2 2 --
     +-- Potomida          
  +--+                     
  |  +-- Pronodularia      
--+                        
  |  +-- Lamprotula_caveata **
  +--+                     
     +-- Lamprotula_leaii **  

Chen et al., 2025 [Publ 4448], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 84 2 2 --
  +-------- Potomida          
--+                           
  |  +----- Pronodularia      
  +--+                        
     |  +-- Lamprotula_caveata **
     +--+                     
        +-- Lamprotula_leaii **  

Wu et al., 2024 [Publ 4350], fig. 6-7

iTtTbiriSoTroTroSiD
2 2 0 2 34 1 1 --
  +----- Potomida          
--+                        
  |  +-- Lamprotula_caveata **
  +--+                     
     +-- Lamprotula_leaii **  

Ouyang et al., 2015 [Publ 4493], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 29 0 0 --
  +-- Lamprotula_caveata **
--+                     
  +-- Lamprotula_leaii **  

Wang et al., 2018 [Publ 3846], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 29 1 1 --
  +----- Potomida          
--+                        
  |  +-- Lamprotula_caveata **
  +--+                     
     +-- Lamprotula_leaii **  

Wu et al., 2024 [Publ 4350], fig. 5b

iTtTbiriSoTroTroSiD
2 2 0 2 34 1 1 --
  +-- Lamprotula_caveata **
--+                     
  +-- Lamprotula_leaii **  

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

iTtTbiriSoTroTroSiD
2 2 0 2 46 2 2 --
  +-------- Potomida          
--+                           
  |  +----- Pronodularia      
  +--+                        
     |  +-- Lamprotula_caveata **
     +--+                     
        +-- Lamprotula_leaii **  

Wu et al., 2021 [Publ 4366], fig. 2_I

iTtTbiriSoTroTroSiD
2 2 0 2 89 2 2 --
  +-------- Potomida          
--+                           
  |  +----- Pronodularia      
  +--+                        
     |  +-- Lamprotula_caveata **
     +--+                     
        +-- Lamprotula_leaii **  

Wu et al., 2024 [Publ 4388], fig. 5

iTtTbiriSoTroTroSiD
2 2 0 2 34 1 1 --
  +----- Potomida          
--+                        
  |  +-- Lamprotula_leaii **  
  +--+                     
     +-- Lamprotula_caveata **

Jeratthitikul et al., 2021 [Publ 4219], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 34 2 2 --
  +----- Potomida        
--+                      
  |  +-- Lamprotula_leaii **
  +--+                   
     +-- Pronodularia    

Konopleva et al., 2023 [Publ 4262], fig. 8

iTtTbiriSoTroTroSiD
1 1 0 1 81 1 1 --
  +-- Lamprotula_leaii **
--+                   
  +-- Potomida        

Wu et al., 2020 [Publ 4121], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 48 2 2 --
  +----- Potomida        
--+                      
  |  +-- Pronodularia    
  +--+                   
     +-- Lamprotula_leaii **

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iTtTbiriSoTroTroSiD
1 1 0 1 75 2 2 --
  +----- Potomida        
--+                      
  |  +-- Lamprotula_leaii **
  +--+                   
     +-- Pronodularia    

Chen et al., 2012 [Publ 3624], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 6 0 0 --
-- Lamprotula_leaii **

Sheng et al., 2014 [Publ 3704], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 7 0 0 --
-- Lamprotula_leaii **

Chen et al., 2025 [Publ 4487], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 40 1 1 --
  +-- Pronodularia    
--+                   
  +-- Lamprotula_leaii **

Chen et al., 2023 [Publ 4302], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 40 1 1 --
  +-- Pronodularia    
--+                   
  +-- Lamprotula_leaii **

Sano et al., 2020 [Publ 4138], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 67 2 2 --
  +----- Lamprotula_leaii **
--+                      
  |  +-- Potomida        
  +--+                   
     +-- Pronodularia    

Zhang et al., 2004 [Publ 2735], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 13 0 0 --
-- Lamprotula_caveata **

Xue et al., 2016 [Publ 3659], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 14 0 0 --
-- Lamprotula_leaii **

Sonowal et al., 2021 [Publ 4221], fig. 3a

iTtTbiriSoTroTroSiD
1 1 0 1 14 1 1 --
  +-- Lamprotula_leaii **
--+                   
  +-- Potomida        

Guerra et al., 2019 [Publ 4168], fig. 1-M

iTtTbiriSoTroTroSiD
1 1 0 1 21 2 2 --
  +----- Potomida        
--+                      
  |  +-- Lamprotula_leaii **
  +--+                   
     +-- Pronodularia    

Bolotov et al., 2018 [Publ 3826], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 81 2 2 --
-- Lamprotula_leaii **

Chen et al., 2025 [Publ 4487], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 40 1 1 --
  +-- Pronodularia    
--+                   
  +-- Lamprotula_leaii **

Huang et al., 2002 [Publ 2657], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 33 0 0 --
-- Lamprotula_leaii **

Wu et al., 2019 [Publ 3864], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 45 2 2 --
  +----- Potomida        
--+                      
  |  +-- Lamprotula_leaii **
  +--+                   
     +-- Pronodularia    

Zhang et al., 2013 [Publ 4214], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 3 0 0 --
-- Lamprotula_leaii **

Sano et al., 2020 [Publ 4138], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 67 2 2 --
  +----- Lamprotula_leaii **
--+                      
  |  +-- Pronodularia    
  +--+                   
     +-- Potomida        

Sonowal et al., 2021 [Publ 4221], fig. 3b

iTtTbiriSoTroTroSiD
1 1 0 1 14 1 1 --
-- Lamprotula_leaii **

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

iTtTbiriSoTroTroSiD
1 1 0 1 38 2 2 --
  +----- Potomida        
--+                      
  |  +-- Lamprotula_leaii **
  +--+                   
     +-- Pronodularia    

Zhou et al., 2016 [Publ 3710], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 19 1 1 --
  +-- Lamprotula_leaii **
--+                   
  +-- Pronodularia    

Smith, 2021 [Publ 4331], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 19 0 0 --
-- Lamprotula_leaii **

Liu et al., 2023 [Publ 4310], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 23 0 0 --
-- Lamprotula_leaii **

Burzynski & Soroka, 2018 [Publ 3862], fig. 5a

iTtTbiriSoTroTroSiD
1 1 0 1 19 2 2 --
  +----- Potomida        
--+                      
  |  +-- Lamprotula_leaii **
  +--+                   
     +-- Pronodularia    

Bolotov et al., 2020 [Publ 4084], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 35 2 2 --
-- Lamprotula_leaii **

Guerra et al., 2019 [Publ 4168], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 40 2 2 --
  +----- Pronodularia    
--+                      
  |  +-- Lamprotula_leaii **
  +--+                   
     +-- Potomida        

Bolotov et al., 2022 [Publ 4250], fig. 8

iTtTbiriSoTroTroSiD
1 1 0 1 48 1 1 --
-- Lamprotula_leaii **

Dai et al., 2025 [Publ 4440], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 44 1 1 --
  +-- Pronodularia    
--+                   
  +-- Lamprotula_leaii **

Chase et al., 2018 [Publ 3861], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 10 0 0 --
-- Lamprotula_leaii **

Burzynski et al., 2017 [Publ 3774], fig. 1-top

iTtTbiriSoTroTroSiD
1 1 0 1 12 2 2 --
  +----- Lamprotula_leaii **
--+                      
  |  +-- Potomida        
  +--+                   
     +-- Pronodularia    

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 36 1 1 --
  +-- Lamprotula_leaii **
--+                   
  +-- Potomida        

Zieritz et al., 2020 [Publ 4102], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 30 0 0 --
-- Lamprotula_leaii **

Aunins et al., 2018 [Publ 3863], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 39 1 1 --
  +-- Lamprotula_caveata **
--+                     
  +-- Potomida          

Jeratthitikul et al., 2021 [Publ 4219], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 33 2 2 --
  +----- Potomida        
--+                      
  |  +-- Pronodularia    
  +--+                   
     +-- Lamprotula_leaii **

Burzynski et al., 2017 [Publ 3774], fig. 1-bottom

iTtTbiriSoTroTroSiD
1 1 0 1 12 2 2 --
  +----- Potomida        
--+                      
  |  +-- Lamprotula_leaii **
  +--+                   
     +-- Pronodularia    

Araujo et al., 2017 [Publ 3801], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 13 2 2 --
  +----- Potomida        
--+                      
  |  +-- Lamprotula_leaii **
  +--+                   
     +-- Pronodularia    

Froufe et al., 2016 [Publ 3727], fig. 3_4

iTtTbiriSoTroTroSiD
1 1 0 1 6 4 2 --
-- Lamprotula_leaii **

Guerra et al., 2019 [Publ 4168], fig. 2-M

iTtTbiriSoTroTroSiD
1 1 0 1 21 2 2 --
  +----- Potomida        
--+                      
  |  +-- Lamprotula_leaii **
  +--+                   
     +-- Pronodularia    

Froufe et al., 2015 [Publ 3664], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 11 2 2 --
  +----- Potomida        
--+                      
  |  +-- Lamprotula_leaii **
  +--+                   
     +-- Pronodularia    

Lopes-Lima et al., 2017 [Publ 3792], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 31 2 2 --
  +----- Potomida        
--+                      
  |  +-- Pronodularia    
  +--+                   
     +-- Lamprotula_leaii **

Chen et al., 2023 [Publ 4302], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 40 1 1 --
  +-- Pronodularia    
--+                   
  +-- Lamprotula_leaii **

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 39 2 2 --
  +----- Potomida          
--+                        
  |  +-- Pronodularia      
  +--+                     
     +-- Lamprotula_caveata **

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iTtTbiriSoTroTroSiD
1 1 0 1 75 2 2 --
  +----- Lamprotula_leaii **
--+                      
  |  +-- Potomida        
  +--+                   
     +-- Pronodularia    

Huang et al., 2018 [Publ 3828], fig. 3_5

iTtTbiriSoTroTroSiD
1 1 0 1 28 1 1 --
  +-- Potomida        
--+                   
  +-- Lamprotula_leaii **

Liu et al., 2023 [Publ 4310], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 23 0 0 --
-- Lamprotula_leaii **

Sano et al., 2020 [Publ 4138], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 67 2 2 --
  +----- Lamprotula_leaii **
--+                      
  |  +-- Pronodularia    
  +--+                   
     +-- Potomida        

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 56 0 0 --
-- Lamprotula_cornuumlunae **

Ouyang et al., 2011 [Publ 3408], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 12 0 0 --
-- Lamprotula_caveata **

Dai et al., 2025 [Publ 4440], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 44 1 1 --
  +-- Pronodularia    
--+                   
  +-- Lamprotula_leaii **

Jeratthitikul et al., 2024 [Publ 4387], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 58 2 2 --
  +----- Pronodularia    
--+                      
  |  +-- Lamprotula_leaii **
  +--+                   
     +-- Potomida        

Wu et al., 2018 [Publ 4143], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 21 1 1 --
  +-- Pronodularia    
--+                   
  +-- Lamprotula_leaii **

Huang et al., 2002 [Publ 2657], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 33 0 0 --
-- Lamprotula_leaii **

Sonowal et al., 2021 [Publ 4221], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 15 1 1 --
-- Lamprotula_leaii **

Wu et al., 2016 [Publ 3711], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 20 1 1 --
  +-- Lamprotula_leaii **
--+                   
  +-- Pronodularia    

Sano et al., 2020 [Publ 4138], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 33 1 1 --
  +-- Lamprotula_caveata **
--+                     
  +-- Pronodularia      

Chase et al., 2018 [Publ 3861], fig. 1-M

iTtTbiriSoTroTroSiD
1 1 0 1 10 0 0 --
-- Lamprotula_leaii **

Sano et al., 2017 [Publ 3799], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 32 1 1 --
  +-- Pronodularia    
--+                   
  +-- Lamprotula_leaii **

Wen et al., 2017 [Publ 3795], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 15 0 0 --
-- Lamprotula_leaii **

Guerra et al., 2019 [Publ 4168], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 40 2 2 --
  +----- Pronodularia    
--+                      
  |  +-- Lamprotula_leaii **
  +--+                   
     +-- Potomida        

Jeratthitikul et al., 2025 [Publ 4423], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 49 2 2 --
  +----- Potomida        
--+                      
  |  +-- Lamprotula_leaii **
  +--+                   
     +-- Pronodularia    

Dai et al., 2024 [Publ 4358], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 52 2 2 --
  +----- Potomida        
--+                      
  |  +-- Lamprotula_leaii **
  +--+                   
     +-- Pronodularia    

Pfeiffer et al., 2021 [Publ 4126], fig. 9

iTtTbiriSoTroTroSiD
1 1 0 1 28 0 0 --
-- Lamprotula_cornuumlunae **

Goncalves et al., 2022 [Publ 4253], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 6 0 0 --
-- Lamprotula_caveata **

Huang et al., 2019 [Publ 3843], fig. 3-M

iTtTbiriSoTroTroSiD
1 1 0 1 22 2 2 --
  +----- Potomida        
--+                      
  |  +-- Lamprotula_leaii **
  +--+                   
     +-- Pronodularia    

Kongim et al., 2023 [Publ 4290], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 14 2 2 --
  +----- Lamprotula_leaii **
--+                      
  |  +-- Pronodularia    
  +--+                   
     +-- Potomida        

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iTtTbiriSoTroTroSiD
1 1 0 1 56 0 0 --
-- Lamprotula_cornuumlunae **

Huang et al., 2018 [Publ 3828], fig. 6-M

iTtTbiriSoTroTroSiD
1 1 0 1 16 2 2 --
  +----- Potomida        
--+                      
  |  +-- Lamprotula_leaii **
  +--+                   
     +-- Pronodularia    

Wu et al., 2023 [Publ 4296], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 22 0 0 --
-- Lamprotula_leaii **

Fonseca et al., 2016 [Publ 3775], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 30 2 2 --
  +----- Potomida        
--+                      
  |  +-- Lamprotula_leaii **
  +--+                   
     +-- Pronodularia    

Konopleva et al., 2021 [Publ 4213], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 25 1 1 --
-- Lamprotula_leaii **

Teiga-Teixeira et al., 2020 [Publ 4173], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 18 0 0 --
-- Lamprotula_leaii **