Bivalvia | Unionida | Unionoidea | Unionidae | Unioninae | Unionini | Oxynaiina

genus Nodularia Conrad, 1853: cladomics

14 Recent valid species (15 total species)

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 14 (15 total)
analyzed in 126 cladograms in 77 publications
maximum number of terminals in an analysis = 11 (mean = 2.73, median = 2)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 73
maximum number of included terminals = 11, (mean = 3.99 median = 3)
proportion of cladograms that support basic monophyly = 0.99

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Nodularia cladogram distances
PCoA Ordination

NJ Tree of Nodularia cladogram distances
NJ Tree

Cladograms

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
11 11 10 12 620 21 6 0 
                          +-- Tchangsinaia         
                    +-----+                        
                    |     +-- Inversiunio          
  +-----------------+                              
  |                 |  +----- Pseudocuneopsis      
  |                 +--+                           
  |                    |  +-- Diaurora             
  |                    +--+                        
--+                       +-- Cuneopsis            
  |                                                
  |  +----------------------- Schistodesmus        
  |  |                                             
  |  |              +-------- Nodularia_dorri__2 **   
  +--+  +-----------+                              
     |  |           |  +----- Nodularia_dorri__1 **   
     |  |           +--+                           
     |  |              |  +-- Nodularia_hanensis **   
     +--+              +--+                        
        |                 +-- Nodularia_micheloti **  
        |                                          
        |  +----------------- Nodularia_breviconcha **
        |  |                                       
        +--+  +-------------- Nodularia_guiensis **   
           |  |                                    
           +--+  +----------- Nodularia_nuxpersicae **
              |  |                                 
              +--+        +-- Nodularia_fusiformans **
                 |  +-----+                        
                 |  |     +-- Nodularia_huana **      
                 +--+                              
                    |  +----- Nodularia_nipponensis **
                    +--+                           
                       |  +-- Nodularia_douglasiae ** 
                       +--+                        
                          +-- Nodularia_dualobtusa **

Hou et al., 2025 [Publ 4401], fig. 4

iTtTbiriSoTroTroSiD
11 11 9 11 17 14 4 9 
  +-------------------------- Pseudocuneopsis      
  |                                                
  |                    +----- Schistodesmus        
--+  +-----------------+                           
  |  |                 |  +-- Tchangsinaia         
  |  |                 +--+                        
  |  |                    +-- Cuneopsis            
  +--+                                             
     |                 +----- Nodularia_dorri **      
     |  +--------------+                           
     |  |              |  +-- Nodularia_hanensis **   
     |  |              +--+                        
     +--+                 +-- Nodularia_micheloti **  
        |                                          
        |  +----------------- Nodularia_breviconcha **
        |  |                                       
        +--+  +-------------- Nodularia_huana **      
           |  |                                    
           +--+  +----------- Nodularia_fusiformans **
              |  |                                 
              +--+  +-------- Nodularia_guiensis **   
                 |  |                              
                 +--+     +-- Nodularia_douglasiae ** 
                    |  +--+                        
                    |  |  +-- Nodularia_dualobtusa ** 
                    +--+                           
                       |  +-- Nodularia_nipponensis **
                       +--+                        
                          +-- Nodularia_nuxpersicae **

Hou et al., 2025 [Publ 4401], fig. 3

iTtTbiriSoTroTroSiD
11 11 9 11 0 0 0 11 
              +----- Nodularia_dorri **      
  +-----------+                           
  |           |  +-- Nodularia_hanensis **   
  |           +--+                        
  |              +-- Nodularia_micheloti **  
--+                                       
  |  +-------------- Nodularia_guiensis **   
  |  |                                    
  |  |           +-- Nodularia_douglasiae ** 
  +--+  +--------+                        
     |  |        +-- Nodularia_dualobtusa ** 
     |  |                                 
     +--+        +-- Nodularia_nipponensis **
        |  +-----+                        
        |  |     +-- Nodularia_nuxpersicae **
        +--+                              
           |  +----- Nodularia_huana **      
           +--+                           
              |  +-- Nodularia_breviconcha **
              +--+                        
                 +-- Nodularia_fusiformans **

Wu et al., 2024 [Publ 4379], fig. 2

iTtTbiriSoTroTroSiD
10 10 8 10 3 3 3 6 
                 +-- Cuneopsis            
                 |                        
  +--------------+-- Tchangsinaia         
  |              |                        
  |              +-- Pseudocuneopsis      
  |                                       
--+           +----- Nodularia_dorri **      
  |  +--------+                           
  |  |        |  +-- Nodularia_hanensis **   
  |  |        +--+                        
  |  |           +-- Nodularia_micheloti **  
  +--+                                    
     |  +----------- Nodularia_breviconcha **
     |  |                                 
     |  |        +-- Nodularia_fusiformans **
     +--+  +-----+                        
        |  |     +-- Nodularia_huana **      
        |  |                              
        +--+     +-- Nodularia_douglasiae ** 
           |  +--+                        
           |  |  +-- Nodularia_dualobtusa ** 
           +--+                           
              |  +-- Nodularia_nuxpersicae **
              +--+                        
                 +-- Nodularia_nipponensis **

Hou et al., 2025 [Publ 4445], fig. 5

iTtTbiriSoTroTroSiD
10 10 8 10 43 12 4 8 
  +----------------------- Pseudocuneopsis      
  |                                             
  |                 +----- Schistodesmus        
--+  +--------------+                           
  |  |              |  +-- Tchangsinaia         
  |  |              +--+                        
  |  |                 +-- Cuneopsis            
  +--+                                          
     |              +----- Nodularia_dorri **      
     |  +-----------+                           
     |  |           |  +-- Nodularia_micheloti **  
     |  |           +--+                        
     +--+              +-- Nodularia_hanensis **   
        |                                       
        |  +-------------- Nodularia_breviconcha **
        |  |                                    
        +--+  +----------- Nodularia_huana **      
           |  |                                 
           +--+  +-------- Nodularia_fusiformans **
              |  |                              
              +--+     +-- Nodularia_nipponensis **
                 |  +--+                        
                 |  |  +-- Nodularia_nuxpersicae **
                 +--+                           
                    |  +-- Nodularia_douglasiae ** 
                    +--+                        
                       +-- Nodularia_dualobtusa **

Hou et al., 2024 [Publ 4400], fig. 4

iTtTbiriSoTroTroSiD
10 10 8 10 66 14 4 8 
  +----------------------- Pseudocuneopsis      
  |                                             
  |                 +----- Schistodesmus        
--+  +--------------+                           
  |  |              |  +-- Tchangsinaia         
  |  |              +--+                        
  |  |                 +-- Cuneopsis            
  +--+                                          
     |              +----- Nodularia_dorri **      
     |  +-----------+                           
     |  |           |  +-- Nodularia_hanensis **   
     |  |           +--+                        
     +--+              +-- Nodularia_micheloti **  
        |                                       
        |  +-------------- Nodularia_breviconcha **
        |  |                                    
        +--+  +----------- Nodularia_huana **      
           |  |                                 
           +--+  +-------- Nodularia_fusiformans **
              |  |                              
              +--+     +-- Nodularia_douglasiae ** 
                 |  +--+                        
                 |  |  +-- Nodularia_dualobtusa ** 
                 +--+                           
                    |  +-- Nodularia_nipponensis **
                    +--+                        
                       +-- Nodularia_nuxpersicae **

Wu et al., 2024 [Publ 4379], fig. 5

iTtTbiriSoTroTroSiD
10 10 8 10 44 9 4 8 
  +----------------------- Pseudocuneopsis      
  |                                             
  |                 +----- Schistodesmus        
--+  +--------------+                           
  |  |              |  +-- Tchangsinaia         
  |  |              +--+                        
  |  |                 +-- Cuneopsis            
  +--+                                          
     |              +----- Nodularia_dorri **      
     |  +-----------+                           
     |  |           |  +-- Nodularia_hanensis **   
     |  |           +--+                        
     +--+              +-- Nodularia_micheloti **  
        |                                       
        |  +-------------- Nodularia_breviconcha **
        |  |                                    
        +--+  +----------- Nodularia_huana **      
           |  |                                 
           +--+  +-------- Nodularia_fusiformans **
              |  |                              
              +--+     +-- Nodularia_douglasiae ** 
                 |  +--+                        
                 |  |  +-- Nodularia_dualobtusa ** 
                 +--+                           
                    |  +-- Nodularia_nipponensis **
                    +--+                        
                       +-- Nodularia_nuxpersicae **

Hou et al., 2025 [Publ 4432], fig. 4

iTtTbiriSoTroTroSiD
10 10 8 10 18 15 4 8 
  +----------------------- Pseudocuneopsis      
  |                                             
  |                 +----- Schistodesmus        
--+  +--------------+                           
  |  |              |  +-- Tchangsinaia         
  |  |              +--+                        
  |  |                 +-- Cuneopsis            
  +--+                                          
     |              +----- Nodularia_dorri **      
     |  +-----------+                           
     |  |           |  +-- Nodularia_hanensis **   
     |  |           +--+                        
     +--+              +-- Nodularia_micheloti **  
        |                                       
        |  +-------------- Nodularia_breviconcha **
        |  |                                    
        +--+  +----------- Nodularia_huana **      
           |  |                                 
           +--+  +-------- Nodularia_fusiformans **
              |  |                              
              +--+     +-- Nodularia_douglasiae ** 
                 |  +--+                        
                 |  |  +-- Nodularia_dualobtusa ** 
                 +--+                           
                    |  +-- Nodularia_nipponensis **
                    +--+                        
                       +-- Nodularia_nuxpersicae **

Wu et al., 2024 [Publ 4379], fig. 4_6

iTtTbiriSoTroTroSiD
10 10 8 10 13 2 1 8 
  +-------------------- Schistodesmus        
  |                                          
  |              +----- Nodularia_dorri **      
--+  +-----------+                           
  |  |           |  +-- Nodularia_micheloti **  
  |  |           +--+                        
  +--+              +-- Nodularia_hanensis **   
     |                                       
     |  +-------------- Nodularia_breviconcha **
     |  |                                    
     +--+  +----------- Nodularia_huana **      
        |  |                                 
        +--+  +-------- Nodularia_fusiformans **
           |  |                              
           +--+     +-- Nodularia_douglasiae ** 
              |  +--+                        
              |  |  +-- Nodularia_dualobtusa ** 
              +--+                           
                 |  +-- Nodularia_nuxpersicae **
                 +--+                        
                    +-- Nodularia_nipponensis **

Choi et al., 2020 [Publ 4169], fig. 2a

iTtTbiriSoTroTroSiD
7 7 6 8 5 4 3 12 
  +----------------- Nodularia_breviconcha **  
  |                                         
--+  +-------------- Nodularia_huana **        
  |  |                                      
  +--+  +----------- Nodularia_fusiformans **  
     |  |                                   
     +--+        +-- Nodularia_nipponensis **  
        |  +-----+                          
        |  |     +-- Nodularia_nuxpersicae **  
        +--+                                
           |  +----- Nodularia_dualobtusa__2 **
           +--+                             
              |  +-- Nodularia_douglasiae **   
              +--+                          
                 +-- Nodularia_dualobtusa__1 **

Choi et al., 2020 [Publ 4169], fig. 5

iTtTbiriSoTroTroSiD
7 7 5 7 3 0 0 9 
  +----------- Nodularia_breviconcha **
  |                                 
  |        +-- Nodularia_fusiformans **
--+  +-----+                        
  |  |     +-- Nodularia_huana **      
  |  |                              
  +--+     +-- Nodularia_dualobtusa ** 
     |  +--+                        
     |  |  +-- Nodularia_douglasiae ** 
     +--+                           
        |  +-- Nodularia_nipponensis **
        +--+                        
           +-- Nodularia_nuxpersicae **

Lopes-Lima et al., 2020 [Publ 4080], fig. 13

iTtTbiriSoTroTroSiD
7 7 5 7 3 0 0 9 
  +----------------- Nodularia_breviconcha **
--+                                       
  |  +-------------- Nodularia_fusiformans **
  +--+                                    
     |  +----------- Nodularia_huana **      
     +--+                                 
        |  +-------- Nodularia_nuxpersicae **
        +--+                              
           |  +----- Nodularia_nipponensis **
           +--+                           
              |  +-- Nodularia_douglasiae ** 
              +--+                        
                 +-- Nodularia_dualobtusa **

Dai et al., 2024 [Publ 4386], fig. 5-BI

iTtTbiriSoTroTroSiD
6 6 4 6 16 3 3 12 
  +----------- Cuneopsis            
  |                                 
  |        +-- Inversiunio          
  +--------+                        
  |        +-- Schistodesmus        
--+                                 
  |        +-- Nodularia_huana **      
  |  +-----+                        
  |  |     +-- Nodularia_breviconcha **
  |  |                              
  +--+     +-- Nodularia_nuxpersicae **
     |  +--+                        
     |  |  +-- Nodularia_nipponensis **
     +--+                           
        |  +-- Nodularia_douglasiae ** 
        +--+                        
           +-- Nodularia_dualobtusa **

Sayenko et al., 2021 [Publ 4189], fig. 6

iTtTbiriSoTroTroSiD
6 6 4 6 7 4 1 8 
  +-------------- Nodularia_breviconcha **
--+                                    
  |  +----------- Nodularia_huana **      
  +--+                                 
     |  +-------- Nodularia_nuxpersicae **
     +--+                              
        |  +----- Nodularia_nipponensis **
        +--+                           
           |  +-- Nodularia_douglasiae ** 
           +--+                        
              +-- Nodularia_dualobtusa **

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
6 6 4 6 116 12 4 8 
              +----- Schistodesmus        
              |                           
  +-----------+----- Cuneopsis            
  |           |                           
  |           |  +-- Tchangsinaia         
--+           +--+                        
  |              +-- Inversiunio          
  |                                       
  |  +-------------- Nodularia_breviconcha **
  +--+                                    
     |  +----------- Nodularia_huana **      
     +--+                                 
        |  +-------- Nodularia_nuxpersicae **
        +--+                              
           |  +----- Nodularia_nipponensis **
           +--+                           
              |  +-- Nodularia_douglasiae ** 
              +--+                        
                 +-- Nodularia_dualobtusa **

Dai et al., 2024 [Publ 4386], fig. 3

iTtTbiriSoTroTroSiD
6 6 4 6 8 2 2 8 
  +----------------- Inversiunio          
  |                                       
--+  +-------------- Cuneopsis            
  |  |                                    
  +--+  +----------- Nodularia_breviconcha **
     |  |                                 
     +--+        +-- Nodularia_fusiformans **
        |  +-----+                        
        |  |     +-- Nodularia_huana **      
        +--+                              
           |  +----- Nodularia_nuxpersicae **
           +--+                           
              |  +-- Nodularia_douglasiae ** 
              +--+                        
                 +-- Nodularia_nipponensis **

Dai et al., 2024 [Publ 4386], fig. 5-ML

iTtTbiriSoTroTroSiD
6 6 3 6 16 3 3 11 
     +----- Cuneopsis            
  +--+                           
  |  |  +-- Inversiunio          
  |  +--+                        
  |     +-- Schistodesmus        
  |                              
--+     +-- Nodularia_nuxpersicae **
  |  +--+                        
  |  |  +-- Nodularia_nipponensis **
  |  |                           
  |  |  +-- Nodularia_huana **      
  +--+--+                        
     |  +-- Nodularia_breviconcha **
     |                           
     |  +-- Nodularia_douglasiae ** 
     +--+                        
        +-- Nodularia_dualobtusa **

Wu et al., 2024 [Publ 4388], fig. 4

iTtTbiriSoTroTroSiD
5 5 3 5 4 0 0 9 
  +-------- Nodularia_breviconcha **
  |                              
--+     +-- Nodularia_douglasiae ** 
  |  +--+                        
  |  |  +-- Nodularia_dualobtusa ** 
  +--+                           
     |  +-- Nodularia_nipponensis **
     +--+                        
        +-- Nodularia_nuxpersicae **

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

iTtTbiriSoTroTroSiD
5 5 3 5 136 16 4 9 
                    +-- Tchangsinaia         
  +-----------------+                        
  |                 +-- Cuneopsis            
--+                                          
  |  +----------------- Pseudocuneopsis      
  +--+                                       
     |  +-------------- Schistodesmus        
     +--+                                    
        |  +----------- Nodularia_breviconcha **
        +--+                                 
           |  +-------- Nodularia_fusiformans **
           +--+                              
              |  +----- Nodularia_nuxpersicae **
              +--+                           
                 |  +-- Nodularia_douglasiae ** 
                 +--+                        
                    +-- Nodularia_dualobtusa **

Dai et al., 2024 [Publ 4386], fig. 6

iTtTbiriSoTroTroSiD
5 5 3 5 37 6 4 11 
  +----------------- Pseudocuneopsis      
  |                                       
  |           +----- Schistodesmus        
--+  +--------+                           
  |  |        |  +-- Tchangsinaia         
  |  |        +--+                        
  +--+           +-- Cuneopsis            
     |                                    
     |  +----------- Nodularia_breviconcha **
     +--+                                 
        |  +-------- Nodularia_fusiformans **
        +--+                              
           |  +----- Nodularia_douglasiae ** 
           +--+                           
              |  +-- Nodularia_nuxpersicae **
              +--+                        
                 +-- Nodularia_nipponensis **

Mabuchi et al., 2021 [Publ 4233], fig. 1

iTtTbiriSoTroTroSiD
4 4 2 4 1 1 1 12 
  +-------- Tchangsinaia         
  |                              
--+     +-- Nodularia **            
  |  +--+                        
  |  |  +-- Nodularia_nipponensis **
  +--+                           
     |  +-- Nodularia_fusiformans **
     +--+                        
        +-- Nodularia_breviconcha **

Wu et al., 2026 [Publ 4811], fig. 4

iTtTbiriSoTroTroSiD
4 4 2 4 91 16 4 10 
  +-------------- Pseudocuneopsis      
  |                                    
  |        +----- Schistodesmus        
--+  +-----+                           
  |  |     |  +-- Tchangsinaia         
  |  |     +--+                        
  +--+        +-- Cuneopsis            
     |                                 
     |  +-------- Nodularia_hanensis **   
     +--+                              
        |  +----- Nodularia_fusiformans **
        +--+                           
           |  +-- Nodularia_douglasiae ** 
           +--+                        
              +-- Nodularia_guiensis **

Liu et al., 2017 [Publ 3939], fig. 2

iTtTbiriSoTroTroSiD
4 4 3 6 11 3 2 10 
  +-------------- Schistodesmus          
  |                                      
  +-------------- Cuneopsis              
--+                                      
  |  +----------- Nodularia_breviconcha **  
  |  |                                   
  +--+  +-------- Nodularia_douglasiae__3 **
     |  |                                
     +--+  +----- Nodularia_douglasiae__2 **
        |  |                             
        +--+  +-- Nodularia_douglasiae__1 **
           |  |                          
           +--+-- Nodularia_nuxpersicae **  
              |                          
              +-- Nodularia_nipponensis **

Choi et al., 2020 [Publ 4169], fig. 3a

iTtTbiriSoTroTroSiD
4 4 3 8 2 1 1 11 
  +----------- Nodularia_breviconcha **   
  |                                    
--+  +-------- Nodularia_nipponensis__3 **
  |  |                                 
  +--+  +----- Nodularia_douglasiae__3 ** 
     |  |                              
     |  |  +-- Nodularia_dualobtusa **    
     +--+  |                           
        |  +-- Nodularia_nipponensis__1 **
        |  |                           
        +--+-- Nodularia_nipponensis__2 **
           |                           
           +-- Nodularia_douglasiae__1 ** 
           |                           
           +-- Nodularia_douglasiae__2 **

Kim et al., 2021 [Publ 4197], fig. 1

iTtTbiriSoTroTroSiD
4 4 2 4 14 3 2 8 
           +-- Cuneopsis            
  +--------+                        
  |        +-- Schistodesmus        
--+                                 
  |  +-------- Nodularia_breviconcha **
  +--+                              
     |  +----- Nodularia_nipponensis **
     +--+                           
        |  +-- Nodularia_dualobtusa ** 
        +--+                        
           +-- Nodularia_douglasiae **

Dai et al., 2025 [Publ 4440], fig. 3

iTtTbiriSoTroTroSiD
4 4 2 4 84 12 4 10 
           +----- Schistodesmus        
  +--------+                           
  |        |  +-- Tchangsinaia         
  |        +--+                        
--+           +-- Cuneopsis            
  |                                    
  |  +----------- Pseudocuneopsis      
  +--+                                 
     |  +-------- Nodularia_breviconcha **
     +--+                              
        |  +----- Nodularia_fusiformans **
        +--+                           
           |  +-- Nodularia_nuxpersicae **
           +--+                        
              +-- Nodularia_douglasiae **

Klishko et al., 2018 [Publ 3945], fig. 8

iTtTbiriSoTroTroSiD
4 4 2 4 4 0 0 8 
  +-------- Nodularia_breviconcha **
--+                              
  |  +----- Nodularia_huana **      
  +--+                           
     |  +-- Nodularia_douglasiae ** 
     +--+                        
        +-- Nodularia_nipponensis **

Wang et al., 2025 [Publ 4488], fig. 3B

iTtTbiriSoTroTroSiD
4 4 2 4 61 19 4 10 
  +-------------- Pseudocuneopsis      
  |                                    
  |        +----- Schistodesmus        
--+  +-----+                           
  |  |     |  +-- Tchangsinaia         
  |  |     +--+                        
  +--+        +-- Cuneopsis            
     |                                 
     |  +-------- Nodularia_hanensis **   
     +--+                              
        |  +----- Nodularia_fusiformans **
        +--+                           
           |  +-- Nodularia_douglasiae ** 
           +--+                        
              +-- Nodularia_guiensis **

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 3 197 9 3 11 
  +----------- Schistodesmus        
  |                                 
--+        +-- Tchangsinaia         
  |  +-----+                        
  |  |     +-- Cuneopsis            
  +--+                              
     |  +----- Nodularia_douglasiae ** 
     +--+                           
        |  +-- Nodularia_huana **      
        +--+                        
           +-- Nodularia_nuxpersicae **

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
3 3 1 3 195 8 3 11 
     +----- Nodularia_douglasiae ** 
  +--+                           
  |  |  +-- Nodularia_huana **      
  |  +--+                        
--+     +-- Nodularia_nuxpersicae **
  |                              
  |  +----- Schistodesmus        
  +--+                           
     |  +-- Tchangsinaia         
     +--+                        
        +-- Cuneopsis

Huang et al., 2019 [Publ 3843], fig. S1

iTtTbiriSoTroTroSiD
3 3 1 3 44 9 3 11 
     +----- Nodularia_huana **      
  +--+                           
  |  |  +-- Nodularia_nuxpersicae **
  |  +--+                        
--+     +-- Nodularia_douglasiae ** 
  |                              
  |  +----- Tchangsinaia         
  +--+                           
     |  +-- Cuneopsis            
     +--+                        
        +-- Schistodesmus

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
3 3 1 3 197 9 3 11 
  +-------- Cuneopsis_celtiformis
  |                              
  +-------- Tchangsinaia         
  |                              
  +-------- Cuneopsis_rufescens  
--+                              
  |     +-- Cuneopsis_szechenyii 
  +-----+                        
  |     +-- Cuneopsis_heudei     
  |                              
  |  +----- Nodularia_huana **      
  +--+                           
     |  +-- Nodularia_douglasiae ** 
     +--+                        
        +-- Nodularia_nuxpersicae **

Fukata & Iigo, 2019 [Publ 4108], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 3 14 5 4 9 
  +-------------- Pseudocuneopsis      
  |                                    
--+           +-- Cuneopsis            
  |  +--------+                        
  |  |        +-- Schistodesmus        
  +--+                                 
     |  +-------- Tchangsinaia         
     +--+                              
        |  +----- Nodularia_nipponensis **
        +--+                           
           |  +-- Nodularia_douglasiae ** 
           +--+                        
              +-- Nodularia_dualobtusa **

Wu et al., 2024 [Publ 4388], fig. 6

iTtTbiriSoTroTroSiD
3 3 1 3 50 4 4 9 
  +----------- Pseudocuneopsis      
  |                                 
  |     +----- Nodularia_breviconcha **
--+  +--+                           
  |  |  |  +-- Nodularia_douglasiae ** 
  |  |  +--+                        
  +--+     +-- Nodularia_nipponensis **
     |                              
     |  +----- Tchangsinaia         
     +--+                           
        |  +-- Cuneopsis            
        +--+                        
           +-- Schistodesmus

Sano et al., 2017 [Publ 3799], fig. 2

iTtTbiriSoTroTroSiD
3 3 2 4 30 5 3 10 
  +----------- Tchangsinaia            
--+                                    
  |  +-------- Nodularia_nipponensis__1 **
  +--+                                 
     |  +----- Nodularia_breviconcha **   
     +--+                              
        |  +-- Nodularia_nipponensis__2 **
        +--+                           
           +-- Nodularia_douglasiae **

Chen et al., 2025 [Publ 4448], fig. 3

iTtTbiriSoTroTroSiD
3 3 1 3 83 16 5 9 
  +----------- Pseudocuneopsis      
  |                                 
  |     +----- Nodularia_breviconcha **
--+  +--+                           
  |  |  |  +-- Nodularia_douglasiae ** 
  |  |  +--+                        
  |  |     +-- Nodularia_nipponensis **
  +--+                              
     |     +-- Tongchuania          
     |  +--+                        
     |  |  +-- Schistodesmus        
     +--+                           
        |  +-- Tchangsinaia         
        +--+                        
           +-- Cuneopsis

Klishko et al., 2019 [Publ 3886], fig. 9

iTtTbiriSoTroTroSiD
3 3 1 3 8 0 0 9 
  +----- Nodularia_breviconcha **
--+                           
  |  +-- Nodularia_douglasiae ** 
  +--+                        
     +-- Nodularia_huana **

Huang et al., 2019 [Publ 3843], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 3 42 9 3 11 
  +----------------- Schistodesmus        
  |                                       
  |           +----- Nodularia_douglasiae ** 
--+  +--------+                           
  |  |        |  +-- Nodularia_huana **      
  |  |        +--+                        
  +--+           +-- Nodularia_nuxpersicae **
     |                                    
     |  +----------- Cuneopsis_rufescens  
     +--+                                 
        |  +-------- Tchangsinaia         
        +--+                              
           |  +----- Cuneopsis_celtiformis
           +--+                           
              |  +-- Cuneopsis_szechenyii 
              +--+                        
                 +-- Cuneopsis_heudei

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iTtTbiriSoTroTroSiD
2 2 0 2 74 4 3 -- 
           +-- Nodularia_douglasiae ** 
  +--------+                        
  |        +-- Nodularia_nuxpersicae **
--+                                 
  |  +-------- Schistodesmus        
  +--+                              
     |  +----- Cuneopsis_szechenyii 
     +--+                           
        |  +-- Tchangsinaia         
        +--+                        
           +-- Cuneopsis_rufescens

Froufe et al., 2016 [Publ 3738], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 12 0 0 -- 
  +-- Nodularia_douglasiae ** 
--+                        
  +-- Nodularia_breviconcha **

Chen et al., 2025 [Publ 4448], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 7 6 1 -- 
  +----- Pseudocuneopsis      
--+                           
  |  +-- Nodularia_nipponensis **
  +--+                        
     +-- Nodularia_douglasiae **

Chen et al., 2025 [Publ 4487], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 39 11 6 -- 
           +-- Tchangsinaia         
  +--------+                        
  |        +-- Inversiunio          
  |                                 
--+        +-- Cuneopsis            
  |  +-----+                        
  |  |     +-- Pseudocuneopsis      
  |  |                              
  +--+     +-- Nodularia_douglasiae ** 
     |  +--+                        
     |  |  +-- Nodularia_breviconcha **
     +--+                           
        |  +-- Diaurora             
        +--+                        
           +-- Schistodesmus

Lopes-Lima et al., 2020 [Publ 4080], fig. 12

iTtTbiriSoTroTroSiD
2 2 0 2 6 4 1 -- 
  +----- Inversiunio          
--+                           
  |  +-- Nodularia_breviconcha **
  +--+                        
     +-- Nodularia_douglasiae **

Chen et al., 2023 [Publ 4302], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 39 12 6 -- 
  +-- Nodularia_douglasiae ** 
--+                        
  +-- Nodularia_breviconcha **

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iTtTbiriSoTroTroSiD
2 2 0 2 94 10 4 -- 
     +----- Pseudocuneopsis      
  +--+                           
  |  |  +-- Nodularia_breviconcha **
  |  +--+                        
--+     +-- Nodularia_douglasiae ** 
  |                              
  |  +----- Tchangsinaia         
  +--+                           
     |  +-- Schistodesmus        
     +--+                        
        +-- Cuneopsis

Sano et al., 2020 [Publ 4138], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 4 66 5 3 -- 
  +-------------- Inversiunio             
  |                                       
--+           +-- Schistodesmus           
  |  +--------+                           
  |  |        +-- Tchangsinaia            
  +--+                                    
     |  +-------- Nodularia_nipponensis__1 **
     +--+                                 
        |  +----- Nodularia_nipponensis__2 **
        +--+                              
           |  +-- Nodularia_nipponensis__3 **
           +--+                           
              +-- Nodularia_douglasiae **

Bolotov et al., 2018 [Publ 3826], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 3 80 0 0 -- 
  +----- Nodularia_douglasiae__1 **
--+                             
  |  +-- Nodularia_nuxpersicae **  
  +--+                          
     +-- Nodularia_douglasiae__2 **

Wu et al., 2021 [Publ 4366], fig. 3-BI

iTtTbiriSoTroTroSiD
2 2 0 2 72 5 4 -- 
  +----------- Pseudocuneopsis      
  |                                 
--+        +-- Nodularia_douglasiae ** 
  |  +-----+                        
  |  |     +-- Nodularia_breviconcha **
  +--+                              
     |  +----- Schistodesmus        
     +--+                           
        |  +-- Cuneopsis            
        +--+                        
           +-- Tchangsinaia

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iTtTbiriSoTroTroSiD
2 2 0 2 90 10 4 -- 
  +-------- Pseudocuneopsis      
  |                              
  |     +-- Nodularia_breviconcha **
--+-----+                        
  |     +-- Nodularia_douglasiae ** 
  |                              
  |  +----- Schistodesmus        
  +--+                           
     |  +-- Tchangsinaia         
     +--+                        
        +-- Cuneopsis

Chen et al., 2025 [Publ 4487], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 39 11 6 -- 
              +-- Tchangsinaia         
  +-----------+                        
  |           +-- Inversiunio          
--+                                    
  |  +----------- Pseudocuneopsis      
  |  |                                 
  +--+  +-------- Cuneopsis            
     |  |                              
     +--+     +-- Nodularia_douglasiae ** 
        |  +--+                        
        |  |  +-- Nodularia_breviconcha **
        +--+                           
           |  +-- Diaurora             
           +--+                        
              +-- Schistodesmus

Lopes-Lima et al., 2020 [Publ 4080], fig. 5

iTtTbiriSoTroTroSiD
2 2 0 2 7 0 0 -- 
  +-- Nodularia_breviconcha **
--+                        
  +-- Nodularia_douglasiae **

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
2 2 0 2 136 4 3 -- 
  +-- Nodularia_douglasiae ** 
--+                        
  +-- Nodularia_nuxpersicae **

Wu et al., 2022 [Publ 4238], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 72 10 4 -- 
  +----------- Pseudocuneopsis      
  |                                 
--+        +-- Nodularia_douglasiae ** 
  |  +-----+                        
  |  |     +-- Nodularia_breviconcha **
  +--+                              
     |  +----- Schistodesmus        
     +--+                           
        |  +-- Tchangsinaia         
        +--+                        
           +-- Cuneopsis

Sano et al., 2020 [Publ 4138], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 3 66 5 3 -- 
  +----------- Inversiunio             
  |                                    
--+        +-- Tchangsinaia            
  |  +-----+                           
  |  |     +-- Schistodesmus           
  +--+                                 
     |  +----- Nodularia_nipponensis__1 **
     +--+                              
        |  +-- Nodularia_nipponensis__2 **
        +--+                           
           +-- Nodularia_douglasiae **

Bolotov et al., 2017 [Publ 3764], fig. S1

iTtTbiriSoTroTroSiD
2 10 0 2 122 4 3 18 
  +-- Unio_crassus         
  |                        
  +-- Unio_pictorum        
  |                        
  +-- Unio_tumidus         
  |                        
  +-- Scriptolamprotula    
  |                        
  +-- Cuneopsis_szechenyii 
--+                        
  +-- Tchangsinaia         
  |                        
  +-- Cuneopsis_rufescens  
  |                        
  +-- Nodularia_douglasiae ** 
  |                        
  +-- Nodularia_nuxpersicae **
  |                        
  +-- Schistodesmus

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iTtTbiriSoTroTroSiD
2 2 0 2 94 10 4 -- 
  +-------------- Pseudocuneopsis          
  |                                        
  |           +-- Nodularia_breviconcha **    
--+  +--------+                            
  |  |        +-- Nodularia_douglasiae **     
  |  |                                     
  +--+        +-- Schistodesmus_lampreyanus
     |  +-----+                            
     |  |     +-- Cuneopsis                
     +--+                                  
        |  +----- Tchangsinaia             
        +--+                               
           |  +-- Schistodesmus_xinyuensis 
           +--+                            
              +-- Schistodesmus_spinosus

Dai et al., 2025 [Publ 4440], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 43 11 6 -- 
  +-- Nodularia_douglasiae ** 
--+                        
  +-- Nodularia_breviconcha **

Klishko et al., 2017 [Publ 3946], fig. 8

iTtTbiriSoTroTroSiD
2 2 0 2 4 0 0 -- 
  +-- Nodularia_douglasiae ** 
--+                        
  +-- Nodularia_breviconcha **

Wu et al., 2021 [Publ 4366], fig. 3-ML

iTtTbiriSoTroTroSiD
2 2 0 2 72 5 4 -- 
  +----------- Pseudocuneopsis      
  |                                 
--+        +-- Nodularia_douglasiae ** 
  |  +-----+                        
  |  |     +-- Nodularia_breviconcha **
  +--+                              
     |  +----- Tchangsinaia         
     +--+                           
        |  +-- Cuneopsis            
        +--+                        
           +-- Schistodesmus

Chen et al., 2023 [Publ 4302], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 39 12 6 -- 
  +-- Nodularia_douglasiae ** 
--+                        
  +-- Nodularia_breviconcha **

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iTtTbiriSoTroTroSiD
2 2 0 2 74 4 3 -- 
           +-- Nodularia_douglasiae ** 
  +--------+                        
  |        +-- Nodularia_nuxpersicae **
--+                                 
  |  +-------- Schistodesmus        
  +--+                              
     |  +----- Cuneopsis_szechenyii 
     +--+                           
        |  +-- Tchangsinaia         
        +--+                        
           +-- Cuneopsis_rufescens

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iTtTbiriSoTroTroSiD
2 2 0 2 90 10 4 -- 
  +-------- Pseudocuneopsis      
  |                              
  |     +-- Nodularia_breviconcha **
--+-----+                        
  |     +-- Nodularia_douglasiae ** 
  |                              
  |  +----- Schistodesmus        
  +--+                           
     |  +-- Tchangsinaia         
     +--+                        
        +-- Cuneopsis

Sano et al., 2020 [Publ 4138], fig. 4

iTtTbiriSoTroTroSiD
2 2 0 4 66 5 3 -- 
        +----- Inversiunio             
  +-----+                              
  |     |  +-- Schistodesmus           
  |     +--+                           
--+        +-- Tchangsinaia            
  |                                    
  |  +-------- Nodularia_nipponensis__3 **
  +--+                                 
     |  +----- Nodularia_nipponensis__1 **
     +--+                              
        |  +-- Nodularia_douglasiae **    
        +--+                           
           +-- Nodularia_nipponensis__2 **

Ouyang et al., 2015 [Publ 4493], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 29 5 3 -- 
        +-- Tchangsinaia         
  +-----+                        
  |     +-- Cuneopsis_rufescens  
  |                              
--+     +-- Cuneopsis_szechenyii 
  |  +--+                        
  |  |  +-- Cuneopsis_celtiformis
  +--+                           
     |  +-- Nodularia_douglasiae ** 
     +--+                        
        +-- Nodularia_huana **

Dai et al., 2025 [Publ 4440], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 43 11 6 -- 
  +-- Nodularia_douglasiae ** 
--+                        
  +-- Nodularia_breviconcha **

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iTtTbiriSoTroTroSiD
2 2 0 2 78 4 3 -- 
        +-- Nodularia_nuxpersicae **
  +-----+                        
  |     +-- Nodularia_douglasiae ** 
--+                              
  |  +----- Schistodesmus        
  +--+                           
     |  +-- Tchangsinaia         
     +--+                        
        +-- Cuneopsis

Wu et al., 2021 [Publ 4366], fig. 4

iTtTbiriSoTroTroSiD
2 2 0 2 67 5 4 -- 
  +----------- Pseudocuneopsis      
  |                                 
--+        +-- Nodularia_douglasiae ** 
  |  +-----+                        
  |  |     +-- Nodularia_breviconcha **
  +--+                              
     |  +----- Schistodesmus        
     +--+                           
        |  +-- Tchangsinaia         
        +--+                        
           +-- Cuneopsis

Sano et al., 2020 [Publ 4138], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 3 32 4 3 -- 
  +----- Cuneopsis               
  |                              
--+  +-- Nodularia_nipponensis__1 **
  |  |                           
  +--+-- Nodularia_douglasiae **    
     |                           
     +-- Nodularia_nipponensis__2 **

Lopes-Lima et al., 2020 [Publ 4080], fig. 11

iTtTbiriSoTroTroSiD
2 2 0 2 10 0 0 -- 
  +-- Nodularia_breviconcha **
--+                        
  +-- Nodularia_douglasiae **

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
2 2 0 2 136 4 3 -- 
        +-- Nodularia_douglasiae ** 
  +-----+                        
  |     +-- Nodularia_nuxpersicae **
--+                              
  |  +----- Cuneopsis            
  +--+                           
     |  +-- Tchangsinaia         
     +--+                        
        +-- Schistodesmus

Dai et al., 2024 [Publ 4358], fig. 4

iTtTbiriSoTroTroSiD
2 2 0 2 51 12 4 -- 
  +----------- Pseudocuneopsis      
  |                                 
--+        +-- Nodularia_breviconcha **
  |  +-----+                        
  |  |     +-- Nodularia_douglasiae ** 
  +--+                              
     |  +----- Schistodesmus        
     +--+                           
        |  +-- Tchangsinaia         
        +--+                        
           +-- Cuneopsis

Ouyang et al., 2015 [Publ 4493], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 29 5 3 -- 
                 +-- Nodularia_douglasiae **    
  +--------------+                           
  |              +-- Nodularia_huana **         
  |                                          
--+           +----- Cuneopsis_rufescens__2  
  |  +--------+                              
  |  |        |  +-- Schistodesmus           
  |  |        +--+                           
  +--+           +-- Cuneopsis_celtiformis__2
     |                                       
     |  +----------- Tchangsinaia            
     +--+                                    
        |  +-------- Cuneopsis_celtiformis__1
        +--+                                 
           |  +----- Cuneopsis_szechenyii__2 
           +--+                              
              |  +-- Cuneopsis_szechenyii__1 
              +--+                           
                 +-- Cuneopsis_rufescens__1

Wu et al., 2022 [Publ 4238], fig. 2d

iTtTbiriSoTroTroSiD
2 2 0 2 37 13 5 -- 
  +-------------- Pseudocuneopsis      
  |                                    
--+           +-- Nodularia_breviconcha **
  |  +--------+                        
  |  |        +-- Nodularia_douglasiae ** 
  +--+                                 
     |  +-------- Schistodesmus        
     +--+                              
        |  +----- Cuneopsis            
        +--+                           
           |  +-- Tchangsinaia         
           +--+                        
              +-- Inversiunio

Wu et al., 2020 [Publ 4121], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 48 4 4 -- 
  +----------- Pseudocuneopsis     
--+                                
  |  +-------- Nodularia_douglasiae **
  +--+                             
     |  +----- Tchangsinaia        
     +--+                          
        |  +-- Cuneopsis           
        +--+                       
           +-- Schistodesmus

Froufe et al., 2019 [Publ 3936], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 43 2 2 -- 
  +----- Nodularia_douglasiae **
--+                          
  |  +-- Tchangsinaia        
  +--+                       
     +-- Schistodesmus

Wu et al., 2018 [Publ 3836], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 52 6 3 -- 
     +-- Nodularia_douglasiae **
  +--+                       
  |  +-- Schistodesmus       
--+                          
  |  +-- Tchangsinaia        
  +--+                       
     +-- Cuneopsis

Lopes-Lima et al., 2024 [Publ 4376], fig. 4-5_MtG

iTtTbiriSoTroTroSiD
1 1 0 1 21 1 1 -- 
  +-- Tchangsinaia        
--+                       
  +-- Nodularia_dualobtusa **

Wu et al., 2021 [Publ 4215], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 23 2 1 -- 
  +-- Nodularia_douglasiae **
--+                       
  +-- Inversiunio

Zhang et al., 2004 [Publ 2735], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 13 6 4 -- 
  +-------------- Tchangsinaia        
--+                                   
  |  +----------- Cuneopsis_heudei    
  +--+                                
     |  +-------- Nodularia_douglasiae **
     +--+                             
        |  +----- Pseudocuneopsis     
        +--+                          
           |  +-- Cuneopsis_rufescens 
           +--+                       
              +-- Schistodesmus

Xue et al., 2016 [Publ 3659], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 14 1 1 -- 
-- Nodularia_douglasiae **

Konopleva et al., 2023 [Publ 4269], fig. S1_4

iTtTbiriSoTroTroSiD
1 1 0 1 55 3 2 -- 
  +----- Nodularia_douglasiae **
--+                          
  |  +-- Tchangsinaia        
  +--+                       
     +-- Cuneopsis

Soroka & Burzynski, 2017 [Publ 3837], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 22 2 2 -- 
  +----- Schistodesmus       
--+                          
  |  +-- Tchangsinaia        
  +--+                       
     +-- Nodularia_douglasiae **

Huang et al., 2002 [Publ 2657], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 33 2 2 -- 
  +-- Nodularia_douglasiae **
--+                       
  +-- Tchangsinaia

Wu et al., 2019 [Publ 3864], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 45 4 4 -- 
  +----------- Pseudocuneopsis     
--+                                
  |  +-------- Nodularia_douglasiae **
  +--+                             
     |  +----- Tchangsinaia        
     +--+                          
        |  +-- Cuneopsis           
        +--+                       
           +-- Schistodesmus

Sano et al., 2017 [Publ 3799], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 20 3 1 -- 
  +-- Nodularia_nipponensis **
--+                        
  +-- Inversiunio

Mabuchi & Nishida, 2020 [Publ 4120], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 11 1 1 -- 
-- Nodularia_douglasiae **

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

iTtTbiriSoTroTroSiD
1 1 0 1 38 3 2 -- 
  +----- Schistodesmus        
--+                           
  |  +-- Nodularia_nipponensis **
  +--+                        
     +-- Tchangsinaia

Froufe et al., 2019 [Publ 3936], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 43 2 2 -- 
  +----- Nodularia_douglasiae **
--+                          
  |  +-- Tchangsinaia        
  +--+                       
     +-- Schistodesmus

Lopes-Lima et al., 2024 [Publ 4376], fig. 4-5_AHE

iTtTbiriSoTroTroSiD
1 1 0 1 21 1 1 -- 
  +-- Tchangsinaia        
--+                       
  +-- Nodularia_douglasiae **

Huang et al., 2019 [Publ 3843], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 55 2 2 -- 
  +----- Nodularia_douglasiae **
--+                          
  |  +-- Tchangsinaia        
  +--+                       
     +-- Schistodesmus

Liu et al., 2023 [Publ 4310], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 23 10 5 -- 
  +----- Cuneopsis           
  |                          
  |  +-- Nodularia_douglasiae **
  +--+                       
--+  +-- Schistodesmus       
  |                          
  |  +-- Tchangsinaia        
  |  |                       
  +--+-- Inversiunio         
     |                       
     +-- Pseudocuneopsis

Wu et al., 2020 [Publ 4147], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 29 5 4 -- 
  +----------- Pseudocuneopsis     
--+                                
  |  +-------- Nodularia_douglasiae **
  +--+                             
     |  +----- Schistodesmus       
     +--+                          
        |  +-- Cuneopsis           
        +--+                       
           +-- Tchangsinaia

Zhou et al., 2007 [Publ 2658], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 17 1 1 -- 
  +-- Nodularia_douglasiae **
--+                       
  +-- Cuneopsis

Wu et al., 2022 [Publ 4238], fig. 2e

iTtTbiriSoTroTroSiD
1 1 0 1 17 8 4 -- 
  +----------- Pseudocuneopsis     
--+                                
  |  +-------- Nodularia_douglasiae **
  +--+                             
     |  +----- Tchangsinaia        
     +--+                          
        |  +-- Cuneopsis           
        +--+                       
           +-- Schistodesmus

Dai et al., 2024 [Publ 4358], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 5 5 2 -- 
  +----- Pseudocuneopsis     
--+                          
  |  +-- Schistodesmus       
  +--+                       
     +-- Nodularia_douglasiae **

Fukata & Iigo, 2020 [Publ 4106], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 53 5 4 -- 
        +-- Cuneopsis           
  +-----+                       
  |     +-- Schistodesmus       
--+                             
  |  +----- Pseudocuneopsis     
  +--+                          
     |  +-- Nodularia_douglasiae **
     +--+                       
        +-- Tchangsinaia

Bolotov et al., 2020 [Publ 4084], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 35 0 0 -- 
-- Nodularia_douglasiae **

Wu et al., 2022 [Publ 4238], fig. 2a_S1

iTtTbiriSoTroTroSiD
1 1 0 1 11 10 5 -- 
  +-------------- Nodularia **      
--+                              
  |  +----------- Pseudocuneopsis
  +--+                           
     |  +-------- Schistodesmus  
     +--+                        
        |  +----- Cuneopsis      
        +--+                     
           |  +-- Tchangsinaia   
           +--+                  
              +-- Inversiunio

Wu et al., 2021 [Publ 4366], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 90 5 3 -- 
  +-------- Nodularia_douglasiae **
--+                             
  |  +----- Cuneopsis           
  +--+                          
     |  +-- Tchangsinaia        
     +--+                       
        +-- Schistodesmus

Cha et al., 2018 [Publ 3916], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 37 2 2 -- 
  +----- Schistodesmus
--+                   
  |  +-- Nodularia **    
  +--+                
     +-- Tchangsinaia

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 36 3 2 -- 
  +----- Schistodesmus        
--+                           
  |  +-- Nodularia_nipponensis **
  +--+                        
     +-- Tchangsinaia

Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11

iTtTbiriSoTroTroSiD
1 1 0 1 24 0 0 -- 
-- Nodularia_douglasiae **

Aunins et al., 2018 [Publ 3863], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 39 3 3 -- 
  +-------- Nodularia_douglasiae **
--+                             
  |  +----- Tchangsinaia        
  +--+                          
     |  +-- Cuneopsis           
     +--+                       
        +-- Schistodesmus

Kano et al., 2019 [Publ 4194], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 9 1 1 -- 
  +-- Inversiunio          
--+                        
  +-- Nodularia_nipponensis **

Wang et al., 2018 [Publ 3865], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 30 1 1 -- 
  +-- Tchangsinaia        
--+                       
  +-- Nodularia_douglasiae **

Araujo et al., 2009 [Publ 2589], fig. 7A

iTtTbiriSoTroTroSiD
1 1 0 1 10 0 0 -- 
-- Nodularia_douglasiae **

Huang et al., 2018 [Publ 3828], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 41 1 1 -- 
  +-- Tchangsinaia        
--+                       
  +-- Nodularia_douglasiae **

Lopes-Lima et al., 2017 [Publ 3792], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 31 2 2 -- 
  +----- Schistodesmus       
--+                          
  |  +-- Tchangsinaia        
  +--+                       
     +-- Nodularia_douglasiae **

Campbell & Lydeard, 2026 [Publ 4810], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 46 8 4 -- 
  +----------- Pseudocuneopsis     
--+                                
  |  +-------- Nodularia_douglasiae **
  +--+                             
     |  +----- Tchangsinaia        
     +--+                          
        |  +-- Cuneopsis           
        +--+                       
           +-- Schistodesmus

Liu et al., 2023 [Publ 4310], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 23 10 5 -- 
  +----------- Cuneopsis           
  |                                
--+        +-- Nodularia_douglasiae **
  |  +-----+                       
  |  |     +-- Schistodesmus       
  +--+                             
     |  +----- Tchangsinaia        
     +--+                          
        |  +-- Inversiunio         
        +--+                       
           +-- Pseudocuneopsis

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 56 1 1 -- 
  +-- Tchangsinaia        
--+                       
  +-- Nodularia_douglasiae **

Dai et al., 2023 [Publ 4285], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 52 4 4 -- 
        +-- Pseudocuneopsis     
  +-----+                       
  |     +-- Nodularia_douglasiae **
--+                             
  |  +----- Schistodesmus       
  +--+                          
     |  +-- Cuneopsis           
     +--+                       
        +-- Tchangsinaia

Ouyang et al., 2011 [Publ 3408], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 12 1 1 -- 
  +-- Nodularia_douglasiae **
--+                       
  +-- Cuneopsis

Wu et al., 2018 [Publ 4143], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 21 0 0 -- 
-- Nodularia_douglasiae **

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 30 0 0 -- 
-- Nodularia_douglasiae **

Huang et al., 2002 [Publ 2657], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 33 2 2 -- 
  +-- Nodularia_douglasiae **
--+                       
  +-- Tchangsinaia

Wu et al., 2016 [Publ 3711], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 20 0 0 -- 
-- Nodularia_douglasiae **

Zhou et al., 2017 [Publ 3788], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 15 1 1 -- 
  +-- Nodularia_douglasiae **
--+                       
  +-- Tchangsinaia

Zhou et al., 2007 [Publ 2658], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 20 3 2 -- 
  +-- Tchangsinaia        
--+                       
  +-- Nodularia_douglasiae **

Wang et al., 2018 [Publ 3846], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 30 2 2 -- 
  +----- Nodularia_douglasiae **
--+                          
  |  +-- Tchangsinaia        
  +--+                       
     +-- Cuneopsis

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

iTtTbiriSoTroTroSiD
1 1 0 1 47 2 2 -- 
  +----- Nodularia_douglasiae **
--+                          
  |  +-- Schistodesmus       
  +--+                       
     +-- Tchangsinaia

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iTtTbiriSoTroTroSiD
1 1 0 1 56 1 1 -- 
  +-- Nodularia_douglasiae **
--+                       
  +-- Tchangsinaia

Wu et al., 2021 [Publ 4366], fig. 2_I

iTtTbiriSoTroTroSiD
1 1 0 1 90 5 3 -- 
  +-------- Nodularia_douglasiae **
--+                             
  |  +----- Cuneopsis           
  +--+                          
     |  +-- Tchangsinaia        
     +--+                       
        +-- Schistodesmus

Wu et al., 2023 [Publ 4296], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 22 9 5 -- 
  +----------- Cuneopsis           
  |                                
--+        +-- Nodularia_douglasiae **
  |  +-----+                       
  |  |     +-- Schistodesmus       
  +--+                             
     |  +----- Tchangsinaia        
     +--+                          
        |  +-- Inversiunio         
        +--+                       
           +-- Pseudocuneopsis

Fonseca et al., 2016 [Publ 3775], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 30 2 2 -- 
  +----- Schistodesmus       
--+                          
  |  +-- Nodularia_douglasiae **
  +--+                       
     +-- Tchangsinaia

Araujo et al., 2018 [Publ 3779], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 17 0 0 -- 
-- Nodularia_douglasiae **